maintainer: Anita Risch

en: ecosystem-coupling grazing vertebrates invertebrates ecosystem-functions exclusion swiss-national-park subalpine grassland ecosystem-multifunctionality

en: Datafile

en:
ID: Plot ID composited of Fence & Treatment number (90 plots total)

Grassland: Name of the six subalpine grasslands where the exclosure setups were erected. In large grasslands (GRIM, MING, STAB), we established four hierarchical exclosure setups (two in short- and two in tall-grass vegetation), whereas in the small grasslands (BOTS, DADA, MARG) only one setup was erected per vegetation type. BOTS = Val dal Botsch, GRIM = Grimmels, MING = Val Mingér, STAB = Alp Stabelchod, DADA = Stabelchod da daint, MARG = Margunet

Fence: Number of exclosure setup (1 - 18). Note that uneven numbers (1,3,5,7,9,11,13,15,17) represent setups established on short-grass vegetation, even numbers (2,4,6,8,10,12,14,16,18) setups established in tall-grass vegetation
Vegetation	Vegetation type, 1 = short-grass vegetation, 2 = tall-grass vegetation; (9 replicate fences on each vegetation type)
Treatment	Progressive animal exclusion, numbers indicate which animals had access to the plot: 1 = Large/medium/small mammals, and invertebrates have access ("L/M/S/I"), 2 = Medium/small mammals, and invertebrates have access (“M/S/I”), 3 = Small mammals and invertebrates have access (“S/I”), 4 = Invertebrates have access (“I”), 5 = No animals have access (“None”)

Bulkden:	Fine fraction soil bulk density (g cm-3)

Soil.moist	: Soil moisture (%)

Soil.org.C: Rhizosphere soil C content (%)

pH: Soil pH

Nema.PC1: First principal component (PC1) of principal component analysis (PCA) of the soil nematode community composition

Nema.PC2: Second principal component (PC2) of principal component analysis (PCA) of the soil nematode community composition

Micro.PC1: 	First principal component (PC1) of principal component analysis (PCA) of the soil microbial community composition

Micro.PC2: Second principal component (PC2) of principal component analysis (PCA) of the soil microbial community composition

Abv.Inv.PC1: First principal component (PC1) of principal component analysis (PCA) of the aboveground-dwelling invertebrate community composition

Abv.Inv.PC2: Second principal component (PC2) of principal component analysis (PCA) of the aboveground-dwelling invertebrate community composition

Arthr.PC1: First principal component (PC1) of principal component analysis (PCA) of the soil arthropod community composition

Arthr.PC2: Second principal component (PC2) of principal component analysis (PCA) of the soil arthropod community composition

Plant.PC1: First principal component (PC1) of principal component analysis (PCA) of the plant community composition

Plant.PC2: Second principal component (PC2) of principal component analysis (PCA) of the plant community composition

Root.biomass: Root biomass (g m-2)

Micro.biomass	: Microbial biomass carbon (mg kg-1)

Plant.richness: Plant species richness

Plant.N: Plant tissue nitrogen content (%)

Soil.N.min: Soil net nitrogen mineralisation (kg ha-1 season)

Soil.resp: Soil carbon dioxide emissions (soil respiration; g m-2 h-1)

en:
This dataset contains all data on which the following publication below is based.

Paper Citation:
 > Risch AC, Ochoa-Hueso R, van der Putten WH, Bump JK, Busse MD, Frey B, Gwiazdowicz DJ, Page-Dumroese DS, Vandegehuchte ML, Zimmermann S, Schütz M. Size-dependent loss of aboveground animals differentially affects grassland ecosystem coupling and functions. 2018. Nature Communications 9: 3684. [doi: 10.1038/s41467-018-06105-4](https://doi.org/10.1038/s41467-018-06105-4).

Please cite this paper together with the citation for the datafile.

#Methods
Study sites
The experimental exclosure setups were installed within the SNP (IUCN category Ia preserve; Dudley 2008), in south-eastern Switzerland. The park covers 172 km2 of forests and subalpine and alpine grasslands along with scattered rock outcrops and scree slopes. The entire area has been protected from human impact (no hunting, fishing, camping or off-trail hiking) since 1914. Large, fairly homogenous patches of short- and tall-grass vegetation, which originate from different historical management and grazing regimes, cover the park’s subalpine grasslands entirely. Short-grass vegetation developed in areas where cattle used to rest (nutrient input) prior to the park’s foundation (14th century to 1914) (Schütz and others 2003, 2006) and is dominated by lawn grass species such as Festuca rubra L., Briza media L. and Agrostis capillaris L. (Schütz and others 2003, 2006). Today, this vegetation type is intensively grazed by diverse vertebrate and invertebrate communities that inhabit the park and consume up to 60% of the available biomass (Risch and others 2013). Tall-grass vegetation developed where cattle formerly grazed, but did not rest, and is dominated by rather nutrient-poor tussocks of Carex sempervirens Vill. and Nardus stricta L. (Schütz and others 2003, 2006). This vegetation type receives considerably less grazing, with only roughly 20% of the biomass consumed (Risch and others 2013). Consequently, the two vegetation types together represent a long-term trajectory of changes in grazing regimes. Underlying bedrock of all grasslands is dolomite, which renders these grasslands rather poor in nutrients regardless of former and current land-use regimes.
Experimental design
To progressively exclude aboveground vertebrate and invertebrate animals, we established 18 size-selective exclosure setups (nine in short-grass, nine in tall-grass vegetation) distributed over six subalpine grasslands across the SNP (Risch and others 2013, 2015). Elevation differences of exclosure locations did not exceed 350 m (between 1975 and 2300 m a.s.l.). The exclosures were established immediately after snowmelt in spring 2009 and were left in place for five consecutive growing seasons (until end of 2013). They were, however, temporarily dismantled every fall (late October after first snowfall) to protect them from avalanches. They were re-established in the same location every spring immediately after snowmelt. Each size-selective exclosure setup consisted of five plots (2 x 3 m) that progressively excluded aboveground vertebrates and invertebrates from large to small. The plots are labelled according to the guilds that had access to them “L/M/S/I”, “M/S/I”, “S/I”, “I”, “None”; L = large mammals, M = medium mammals, S = small mammals, I = invertebrates, None = no animals had access. As we only had permission to have the experimental setup in place for five consecutive growing seasons, the experiment had to be completely dismantled in the late fall of 2013 and all material removed from the SNP.
Our exclosure design was aimed at excluding mammalian herbivores, but naturally also excluded the few medium and small mammalian predators, as well as the entire aboveground invertebrate food web. A total of 26 large to small mammal species can be found in the SNP, but large apex predators are missing (wolf, bear, lynx) . Reptiles, amphibians and birds are scarce to absent in the subalpine grasslands under study. Only two reptile species occur in the park and they are confined to rocky areas that warm up enough for them to survive. One frog species spawns in an isolated pond far from our grasslands. Only three bird species occasionally feed on the subalpine grasslands. Using game cameras (Moultrie 6MP Game Spy I-60 Infrared Digital Game Camera, Moultrie Feeders, Alabaster, AL, USA), we did observe that the medium- and small-sized mammals (marmot/hares and mice) were not afraid to enter the fences and feed on their designated plots. We never spotted reptiles, amphibians or birds on camera. We distinguished between 59 higher aboveground-dwelling invertebrate taxa that our size-selective exclosures excluded (see also methods for aboveground-dwelling invertebrates below).
The “L/M/S/I” plot (not fenced) was located at least 5 m from the 2.1 m tall and 7 x 9 m large main electrical fence that enclosed the other four plots. The bottom wire of this fence was mounted at 0.5 m height and was not electrified to enable safe access for medium and small mammals, while fencing out the large ones. Within each main fence, we randomly established four 2 x 3 m plots separated by 1-m wide walkways from one another and from the main fence line: 1) the “M/S/I” plots were unfenced, allowing access to all but the large mammals; 2) the “S/I” plots (10 x 10 cm electrical mesh fence) excluded all medium-sized mammals. Note that the bottom 10 cm of this fence remained non-electrified to enable safe access for small mammals; 3) the “I” plots (2 x 2 cm metal mesh fence) excluded all mammals. We double-folded the mesh at the bottom 50 cm to reduce the mesh size to smaller than 1 x 1 cm openings; and 4) the “None” plots were surrounded by a 1 m tall mosquito net (1.5 x 2 mm) to exclude all animals. The top of the plot was covered with a mosquito-meshed wooden frame mounted to the corner posts (roof). We treated these plots a few times with biocompatible insecticide (Clean kill original, Eco Belle GmbH, Waldshut-Tiengen, Germany) to remove insects that might have entered during data collection or that hatched from the soil, but amounts were negligible and did not impact soil moisture conditions within these plots.
To assess whether the design of the “None” exclosure (mesh and roof) affected the response variables within the plots and, therefore, influenced the results, we established an additional six “micro-climate control” exclosures (one in each of the six grasslands) (Risch and others 2013, 2015). These exclosures were built as the “None” exclosures but were open at the bottom (20 cm) of the 3-m side of the fence facing away from the prevailing wind direction to allow invertebrates to enter. A 20-cm high and 3-m long strip of metal mesh was used to block access to small mammals. Thus, this construction allowed a comparable micro-climate to the “None” plots, but also a comparable feeding pressure by invertebrates to the “I” plots. We compared various properties within these exclosures against one another to assess if our construction altered the conditions in the “None” plots. We showed that differences in plant (e.g., vegetation height, aboveground biomass) and soil properties (e.g., soil temperature, moisture) found between the “I” and the “None” treatments were not due to the construction of the “None” exclosure, but a function of animal exclusions, although the amount of UV light reaching the plant canopy was significantly reduced (Risch and others 2013).
Aboveground invertebrate sampling
Aboveground invertebrates were sampled with two different methods to capture both ground- and plant-dwelling organisms: 1) we randomly placed two pitfall traps (67 mm in diameter, covered with a roof) filled with 20% propylene glycol in one 1 x 1 m subplot of the 2 x 3 m treatment plots in spring 2013 (May) and emptied them every two weeks until late September 2013 (Vandegehuchte and others 2017b, 2018). A pitfall trap consisted of a plastic cylinder (13 cm depth, 6.75 cm diameter). Within each cylinder we placed a 100 ml plastic vial with outer diameter 6.70 cm and on top of the cylinder we placed a plastic funnel to guide the invertebrates into the vials. Each trap was cover with a cone-shaped and transparent plastic roof to protect the trap from rain (Vandegehuchte and others 2017b, 2018). Note that in the “None” plots only one trap was placed as control to check for effectiveness of the exclosure. 2) We vacuumed all invertebrates from a 60 x 60 cm area on another 1 x 1 m subplot with a suction sampler (Vortis, Burkhard manufacturing CO, Ltd., Rickmansworth, Hertfordshire, UK) every month from June to September 2013 (Vandegehuchte and others 2017b, 2018). For this purpose, we quickly placed a square plastic frame (60 x 60 x 40 cm) with a closable mosquito mesh sleeve attached to the top edge into the plot from the outside. The suction sample was then inserted into through the sleeve and operated for 45 s to collect the invertebrates (Vandegehuchte and others 2017b, 2018).
We sorted the ≈100 000 individuals collected with both methods by hand and identified each individual morphologically to the lowest taxonomic level feasible (59 taxa, including orders, suborders, subfamilies, families; phylum for Mollusca). These taxa belonged to the following feeding types: 19 herbivores, 16 detritivores, 9 predators, 8 mixed feeders, 5 omnivores and 2 non-classified feeders (or not feeding as adults) (Vandegehuchte and others 2017b). We summed the numbers from the two pitfall traps and the suction sampling over the course of the 2013 season to represent the aboveground invertebrate abundance and community composition of a plot. Note: we did not specifically attempt to catch flying invertebrates with e.g., sticky traps, thus a few flying insects may have been missed with our vacuum sampling approach.
Sampling of plant properties
The vascular plant species composition was assessed at peak biomass every summer (July) by estimating the frequency of occurrence of each species with the pin count method in each plot (Frank and McNaughton 1990). A total of 172 taxa occurred within our 90 plots and we calculated plant species richness for each plot separately. We used the 2013 data in this study. Plant quality was assessed every year in July and September; here we use plant quality at the end of the experiment (September 2013). Two 10 x 100 cm wide strips of vegetation per plot were clipped, combined, dried at 65°C, and ground (Pulverisette 16, Fritsch, Idar-Oberstein, Germany) to pass through a 0.5 mm sieve. Twenty randomly selected samples across all treatments were analysed for N (Leco TruSpec Analyser, Leco, St. Joseph, Michigan, USA) (Vandegehuchte and others 2015). Nitrogen concentrations of the other samples were then estimated from models established for the experiment and the entire SNP relating Fourier transform-near infrared reflectance (FT-NIR) spectra to the measured values of N using a multi-purpose FT-NIR spectrometer (Bruker Optics, Fällanden, Switzerland) (Vandegehuchte and others 2015). Root biomass was sampled every fall by collecting five 2.2 cm diameter x 10 cm deep soil samples (Giddings Machine Company, Windsor, CO, USA) per plot (450 samples year-1). The samples were dried at 30 °C and roots were sorted from the sample by hand. We sorted each sample for 1 h which allowed to retrieve over 90% of all roots present in the samples (Risch and others 2013). The roots were then dried at 65 °C for 48 and weighed to the nearest mg. We averaged the values per plot and used the 2013 data only in this study.
Sampling of edaphic communities
In 2009, 2010, and 2011 we collected three composited soil samples (5 cm diameter x 10 cm depth; AMS Samplers, American Falls, ID, USA) and assessed bacterial community structure using T-RFLP profiling (Liu and others 1997; Blackwood and others 2003; Hodel and others 2014). We detected a total of 89 operational taxonomic units (OTUs). These values are in accordance with other studies reporting OTU richness (Wirthner and others 2011; Zumsteg and others 2012; Meola and others 2014) using T-RFLP profiling, a method that detects the most abundant, and thus likely, the most relevant, taxa. We averaged the data over the three years of collections for our calculations. Microbial biomass carbon (MBC) was determined with the substrate-induced method (Anderson and Domsch 1978) every fall (September) between 2009 and 2013 by collecting three mineral soil samples (5 cm diameter × 10 cm mineral soil core, AMS Samplers, American Falls, ID, USA). The three samples were combined (90 samples for each sampling year), immediately put on ice, taken to the laboratory, passed through a 2-mm sieve and stored at 4°C. Again, we only used the 2013 data in this study.
Soil samples (5 cm diameter x 10 cm depth) to extract soil arthropods were collected in June, July, and August 2011 with a soil corer lined with a plastic sleeve to ensure an undisturbed sample (total of 270 samples). The plastic line core was immediately sealed on both ends using cling film and put into a cooler. All plots were sampled within three days and the extraction of arthropods started the evening of the sampling day using a high-gradient Tullgren funnel apparatus (Crossley and Blair 1991; Vandegehuchte and others 2015). Samples were kept in the extractor for four days and the soil arthropods were collected in 95% ethanol. All individuals were counted and each individual was identified morphologically to the lowest level feasible [76 taxa, including orders, suborders, subfamilies, families (Protura, Thysanoptera, Aphidina, Psylina, Coleoptera, Brachycera, Nematocera, Auchenorryncha, Heteroptera, Formicidae); sub-phylum for Myriapoda, for Acari and Collembola also including morpho-species). Note that we also included larval stages (nine of the 76 taxa) (Vandegehuchte and others 2015). All data were summed over the season. A detailed species list for mites and collembolans is published (Vandegehuchte and others 2017a) [https://doi.org/10.1371/journal.pone.0118679.s001]. Earthworms are rare in the SNP and therefore were not included. We collected eight random 2.2 cm diameter x 10 cm deep soil cores from each plot in September 2013 to determine the soil nematode community composition. The samples were mixed and the nematodes were extracted from 100 ml of fresh soil using Oostenbrink elutriators (Oostenbrink 1960). All nematodes in a 1 ml of the 10 ml extract were counted, a minimum of 150 individuals sample-1 were identified to genus or family level using (Bongers 1988), the numbers of all nematodes were extrapolated to the entire sample and expressed for a 100 g dry sample. In total we identified 63 genus or family levels (Vandegehuchte and others 2015). The list of all the nematodes found is published (Vandegehuchte and others 2015) [http://www.oikosjournal.org/appendix/oik-03341] or DOI: [doi: 10.1111/oik.03341].
We are aware that sampling soil microbes from 2009 to 2011 and soil arthropods in 2011 was not ideal, but we are positive that this does not bias the results. Most of the parameters measured in our experiment either already showed a treatment response after the first growing season (e.g., plant biomass) or did not respond over the entire time experiment (e.g., microbial biomass C). The microbial community composition (2009 – 2011) was highly influenced by inter-annual differences in temperature and precipitation, but did not differ between treatments or vegetation types (Hodel and others 2014). We therefore felt comfortable using the 2009 through 2011 data for describing the soil microbial community in our experimental treatments. Similarly, we are positive that our soil arthropod data are representative. We did assess soil arthropods in August 2012 and found no differences to the August 2011 data. However, we did not feel comfortable combining the 2011 June, July, August data with only August data for 2012 for our analyses.
Sampling of soil properties
We collected three soil samples (5 cm diameter x 10 cm depth) in each plot in September 2013 after removing the vegetation. First, we collected the top layer of mineral soil rich in organic matter, the surface organic layer or rhizosphere, typically 1 to 3 cm in depth with a soil corer (AMS Samples, American Falls, Idaho, USA). Second, we collected a 10 cm mineral soil core beneath this surface layer. The cores for each layer were composited, dried at 65 °C for 48 h and fine-ground to pass a 0.5 mm screen. We then analysed all samples for total C using a Leco TruSpec Analyser (Leco, St. Joseph, Michigan, USA). Mineral soil pH was measured potentiometrically in 1:2 soil:CaCl2 solution with an equilibration time of 30 min.
Soil net N mineralisation was assessed during the 2013 growing season (Risch and others 2015). For this purpose, we randomly collected a 5 cm diameter x10 cm deep soil sample with a soil corer (AMS Samples, American Falls, Idaho, USA) after clipping the vegetation in June 2013. After weighing and sieving (4 mm mesh) the soil, we extracted a 20 g subsample in 1 mol l-1 KCl for 1.5 h on an end-over-end shaker and thereafter filtered it through ashless folded filter paper (DF 5895 150, ALBET LabScience, Hahnenmühle FineArt GmbH, Dassel, Germany). From these filtrates NO3- concentrations were measured colorimetrically (Norman and Stucki 1981) and NH4+with flow injection analysis (FIAS 300, Perkin Elmer, Waltham Massachusetts, USA) (Risch and others 2015). We dried the rest of the sample 105 °C to constant mass to determine fine,fraction bulk density. A second soil sample was collected within each plot in June 2013 with a corer lined with a 5 x 13 cm aluminium cylinder. The corer was driven 11.5 cm deep into the soil so that the top 1.5 cm of the cylinder remained empty. Into this space we placed a polyester bag (250 µm) filled an ion-exchanger resin to capture the incoming N. The bag was filled with a 1:1 mixture of acidic and alkaline exchanger resin (ion-exchanger I KA/ion exchanger IIIAA, Merck AG, Darmstadt, Germany). We then removed 1.5 cm soil at the bottom of the cylinder and placed a second resin exchanger bag into this space to capture the N leached from the soil column. To assure that the exchange resin was saturated with H+ and Cl- prior to filling the bags, the mixture was stirred with 1.2 ml l-1 HCl for 1 h and then rinsed with demineralized water until the electrical conductivity of the water reached 5 µm cm-1. The cylinder with the resin bags in place was reinserted into the soil with the top flush to the soil surface and incubated for three months. We recollected the cylinders in September 2013. Each resin bag and 20 g of sieved soil (4 mm mesh) from each cylinder were then separately extracted with KCl and NO3- and NH4+ concentrations were measured. Nitrate and NH4+ concentrations of all samples were then converted to a content basis by multiplying their values with fine fraction bulk density. Net N mineralisation was thereafter calculated as the difference between the N content of the samples collected at the end of the three-month incubation (including the N extracted from the bottom resin bag) and the N content at the beginning of the incubation (Risch and others 2015).
Soil CO2 emissions were measured every two weeks between 0900 and 1700 hrs from early May through late September 2013 with a PP-Systems SRC-1 soil respiration chamber (15 cm high, 10 cm diameter; closed circuit) attached to a PP-Systems EGM-4 infrared gas analyser (PP-Systems, Amesbury, MA, USA) on two locations per plot (Risch and others 2013). The chamber was placed on randomly placed, permanently installed PVC collars (10 cm diameter) driven 5 cm into the soil at the beginning of the study (Risch and others 2013). Freshly germinated plants growing within the collars were removed prior to each measurement to avoid measuring plant respiration or photosynthesis. The two measurements collected per plot and sampling date were averaged.
Soil moisture (with time domain reflectometry; Field-Scout TDR-100, Spectrum Technologies, Plainfield, Illionois, USA) and temperature (with a waterproof digital pocket thermometer; Barnstead International, Dubuque, Iowa, USA) were measured at five random locations per plot every two weeks during the growing seasons during the experiment for the 0 to 10 cm depth (Risch and others 2013, 2015). As soil moisture and soil temperature were highly negatively correlated (Risch and others 2013), we only used soil moisture for this study. We used plot-level averages of all values available to capture soil moisture variability during the five years of the experiment. The results remained unchanged when we only used soil moisture from the 2013 growing season.
Numeral calculations and statistical analyses
Ecosystem coupling. We conducted principal component analyses (PCAs; unscaled) at the complete dataset level using the abundances of each taxonomical entity to describe each of the five different communities used in this study: aboveground-dwelling invertebrates, vascular plants, soil microorganisms, soil arthropods and soil nematodes. We retained the first two components (PCA axis 1 and PCA axis 2) of each analysis as we found them to adequately represent the temporal and spatial variability of our 90 treatment plots in previous studies55,67. Together they explained a total of 71.70% of the variation for aboveground invertebrates, 44.36% for plants, 44.85% for soil microorganisms, 61.85% for soil arthropods and 77.19% for soil nematodes. In addition, we used soil pH and soil organic C content as a proxy for soil chemical properties, soil bulk density as a proxy for soil physical properties and soil moisture (negatively correlated with soil temperature) as a proxy for soil micro-climatic conditions for an overall total of fourteen constituents.
We calculated ecosystem coupling9 for each exclosure treatment within each vegetation type (i.e., 2  5 treatment combinations in total) as an integrated measure of pairwise ecological interactions between ecosystem constituents representing ecological communities and the soil abiotic environment. These ecological interactions are defined by non-parametric Spearman rank correlation analyses between two constituents, excluding interactions involving two abiotic constituents (e.g., soil pH vs. soil moisture) and interactions between the first (PC1) and second (PC2) component of each community type, as these are orthogonal by definition. Interactions between abiotic constituents were excluded from the analyses because the focus of our study was on communities and how they interact with one another and their surrounding environment; therefore, including abiotic-abiotic interactions was not of interest here. Given that the effectiveness of our experimental design resulted in that no community composition data of aboveground-dwelling invertebrates was available for the “None” plots (all animals excluded), only thirteen instead of fourteen constituents were included in the ecosystem coupling calculations for this treatment. The complete absence of aboveground invertebrates represents the most extreme case of disturbance between aboveground animal communities and the rest of the ecosystem constituents. This may have resulted in a slight overestimation of ecosystem coupling for these plots.
	Average ecosystem coupling was calculated as follows:
Ecosystem coupling=
where Xi is the absolute Coupling was calculated value of the Spearman’s rho coefficient of the ith correlation for each treatment within each vegetation type (i.e., based on nine replicates each), considering and n is the number of pairwise comparisons considered (n = a total of 80; interactions (56 in the case of the “None” treatment). We considered a total of 40 biotic-biotic interactions (i.e., concerning two community-level principal components such as plants and microbes; 24 in the case of the “None” treatment) and 40 abiotic-biotic (i.e., concerning one community-level principal component and one abiotic factor, e.g., plant community and soil properties; 32 in the case of the “None” treatment).	Coupling was calculated for each treatment within each vegetation type (i.e., based on nine replicates each), considering a total of 80 interactions (56 in the case of the “None” treatment). We considered a total of 40 biotic-biotic interactions (i.e., concerning two community-level principal components such as plants and microbes; 24 in the case of the “None” treatment) and 40 abiotic-biotic (i.e., concerning one community-level principal component and one abiotic factor, e.g., plant community and soil properties; 32 in the case of the “None” treatment).
To establish whether constituents were significantly and positively coupled within treatments (i.e., the average of their correlation coefficients were greater than in a null model where correlation only happens by chance), we calculated one-tailed p-values based on permutation tests with 999 permutations.

We considered six ecosystem functions and process rates commonly used to assess ecosystem functioning (Meyer and others 2015; Manning and others 2018). Plant N content represents a measure of forage quality, while plant richness has been shown to stabilise biomass production, thus allowing the system to respond to changes in herbivory. Soil net N mineralisation, soil respiration, root biomass, and microbial biomass represent fluxes or stocks of energy. For all functions and processes higher values represent higher functioning (Manning and others 2018). All these variables were measured in the last year of the experiment (2013). We then quantified ecosystem multifunctionality using the multiple threshold approach (Byrnes and others 2014; Manning and others 2018), which considers the number of functions that are above a certain threshold, over a series of threshold values (typically 10-99%) that are defined based on the maximum value of each function. We weighted all our functions equally for these calculations (Manning and others 2018). The number of functions in a plot with values higher than a given threshold value for the respective function is summed up. The sum represents ecosystem multifunctionality for that plot. Given that choosing any particular threshold as a measure of ecosystem multifunctionality is arbitrary, we calculated the average of thresholds from 10-90% (in 10% intervals) as a more integrated representation of ecosystem multifunctionality.
We used Pearson correlations to explore the relationships between ecosystem coupling (all interactions, biotic-biotic interactions, abiotic-biotic interactions involving above- and belowground constituents, and all interactions, biotic-biotic interactions, abiotic-biotic interactions involving belowground constituents only) and ecosystem multifunctionality by calculating the slopes of all relationships between ecosystem coupling and multifunctionality for all thresholds between 10 and 99%. We also related ecosystem coupling with the average of multifunctionality at thresholds between 30-80% as explained before and considered this correlation as a robust indication of the type of association between these two variables. In addition, we explored the relationships between ecosystem coupling (all interactions, biotic-biotic interactions, abiotic-biotic interactions involving above- and belowground constituents, and all interactions, biotic-biotic interactions, abiotic-biotic interactions involving belowground constituents only) and individual ecosystem functions. The effects of exclosures and vegetation type on individual functions and multifunctionality were evaluated using linear mixed effects models ('lme' function of the nlme package), with exclosure and vegetation type as fixed effects and fence as a random factor. All statistical analyses and numerical calculations were done in R version 3.4.0 (R Core Team 2016).

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- R Core Team. 2016. R: A language and environment for statistical computing. Vienna, Austria: R Foundation for Statistical Computing
- Risch AC, Haynes AG, Busse MD, Filli F, Schütz M. 2013. The response of soil CO2 fluxes to progressively excluding vertebrate and invertebrate herbivores depends on ecosystem type. Ecosystems 16:1192–202.
- Risch AC, Schütz M, Vandegehuchte ML, Van Der Putten WH, Duyts H, Raschein U, Gwiazdowicz DJ, Busse MD, Page-Dumroese DS, Zimmermann S. 2015. Aboveground vertebrate and invertebrate herbivore impact on net N mineralization in subalpine grasslands. Ecology 96:3312–22.
- Schütz M, Risch AC, Achermann G, Thiel-Egenter C, Page-Dumroese DS, Jurgensen MF, Edwards PJ. 2006. Phosphorus translocation by red deer on a subalpine grassland in the Central European Alps. Ecosystems 9:624–633.
- Schütz M, Risch AC, Leuzinger E, Krüsi BO, Achermann G. 2003. Impact of herbivory by red deer (Cervus elaphus L.) on patterns and processes in subalpine grasslands in the Swiss National Park. For Ecol Manage 181:177–88.
- Vandegehuchte ML, van der Putten WH, Duyts H, Schütz M, Risch AC. 2017a. Aboveground mammal and invertebrate exclusions cause consistent changes in soil food webs of two subalpine grassland types, but mechanisms are system-speciﬁc. Oikos 126:212–23.
- Vandegehuchte ML, Raschein U, Schütz M, Gwiazdowicz DJ, Risch AC. 2015. Indirect short- and long-term effects of aboveground invertebrate and vertebrate herbivores on soil microarthropod communities. PLoS One 10:e0118679.
- Vandegehuchte ML, Schütz M, de Schaetzen F, Risch AC. 2017b. Mammal-induced trophic cascades in invertebrate food webs are modulated by grazing intensity in subalpine grassland. J Anim Ecol 86:1434–46.
- Vandegehuchte ML, Trivellone V, Schütz M, Firn J, de Schaetzen F, Risch AC. 2018. Mammalian herbivores affect leafhoppers associated with specific plant functional types at different timescales. Funct Ecol 32:545–55.
- Wirthner S, Frey B, Busse MD, Schütz M, Risch AC. 2011. Effects of wild boar (Sus scrofa L.) rooting on the bacterial community structure in mixed-hardwood forest soils in Switzerland. Eur J Soil Biol 47:296–302. http://dx.doi.org/10.1016/j.ejsobi.2011.07.003
- Zumsteg A, Luster J, Göransson H, Smittenberg RH, Brunner I, Bernasconi SM, Zeyer J, Frey B. 2012. Bacterial, Archaeal and Fungal Succession in the Forefield of a Receding Glacier. Microb Ecol 63:552–64. https://doi.org/10.1007/s00248-011-9991-8

ecosystem-coupling

display_name: ecosystem-coupling

ecosystem-functions

display_name: ecosystem-functions

ecosystem-multifunctionality

display_name: ecosystem-multifunctionality

exclusion

display_name: exclusion

grassland

display_name: grassland

grazing

display_name: grazing

invertebrates

display_name: invertebrates

subalpine

display_name: subalpine

swiss-national-park

display_name: swiss-national-park

vertebrates

display_name: vertebrates

en: Ecosystem coupling and multifunctionality - exclosure experiment


maintainer: EnviDat Support

en: topsoil-removal fungi soil-chemistry soil nematodes bacteria biodiversity soil-biology soil-biodiversity plants restoration grassland

state: active

type: dataset

en: Ecosystem coupling_interactions

media_type: XLSX

en:
Treatment	Treatment groups; Y.4-18 = restored grasslands; Initial = intensively managed grasslands; Target = species-rich semi-wet to semi-dry grasslands
Type	Interactions type; A.A = abiotic.abiotic; A.B = abiotic.biotic; B.B = biotic.biotic
Correlation	non-parametric Spearman rank correlations between biotic and abiotic parameters (ecological interactions)

format: XLS

en: Distance to target results

en:
Community	biotic community (vegetation, nematoda, fungi, prokaryota)
Treatment	Treatment groups; Y.4-30= restored grasslands (time since restoration); Initial=intensively managed grasslands (I)
Comparison	within- or between-community Bray-Curtis dissimilarity distance comparison
Label	within-community distances of initial/restored grasslands: treatment abbreviation; between-communities distances of initial/restored-to-target grasslands: treatment abbreviation plus Target (T; species-rich grasslands)
mean	Mean distance to Target (6 plots per treatment group)
sd	Standard deviation of distances to Target (6 plots per treatment group)

en: Fungi data

en:
Sample.data
Plot.ID	consecutive numbers for plots (in total 36 plots)
Treatment	One restoration treatment and two reference treatments. Topsoil+Propagules= Topsoil removal (upper 10-30 cm) combined with introduction of target plant species; Initial=intensively managed grasslands; Target= species-rich semi-wet to semi-dry grasslands
treat.yr	Treatment abbreviation. For restored grasslands: Treatment abbreviation + time since restoration; Initial + Target = NA
yr.trt	year when topsoil was removed; Initial + Target = NA
Age	time since restoration (sampling year 2018 - yr.trt); Initial + Target = NA
Age.class	restored sites of similar age grouped into four age classes. Y.4 = Year 3+4 (recently restored); Y.18 = Year 17+19 (mid-term); Y.24 = Year 23+25 (long-term); Y.30 = Year 27+32 (long-term); Initial + Target = NA
Cluster	clustered spatial arrangment of sites in the field (see Supporting Information Fig. S1)
Site	clustered spatial arrangment (replicates) of plots per site (see Supporting Information Fig. S1)

Taxonomy
OTU.ID	consecutive numbers for operational taxonomic units (OTUe 1-7794)
Taxonomic rank	Domain, Phylum, Class, Order, Family, Genus, Species

Abundance
columns	Plot.ID (see Sample.data)
rows	OTU.ID (see Taxonomy)

en: Vegetation data

en:
Plot.ID	consecutive numbers for plots (in total 36 plots)
Treatment	One restoration treatment and two reference treatments. Topsoil+Propagules= Topsoil removal (upper 10-30 cm) combined with introduction of target plant species; Initial=intensively managed grasslands; Target= species-rich semi-wet to semi-dry grasslands
treat.yr	Treatment abbreviation. For restored grasslands: Treatment abbreviation + time since restoration; Initial + Target = NA
yr.trt	year when topsoil was removed; Initial + Target = NA
Age	time since restoration (sampling year 2018 - yr.trt); Initial + Target = NA
Age.class	restored sites of similar age grouped into four age classes. Y.4 = Year 3+4 (recently restored); Y.18 = Year 17+19 (mid-term); Y.24 = Year 23+25 (long-term); Y.30 = Year 27+32 (long-term); Initial + Target = NA
Cluster	clustered spatial arrangment of sites in the field (see Supporting Information Fig. S1)
Site	clustered spatial arrangment (replicates) of plots per site (see Supporting Information Fig. S1)
column I-LW	Plant abundance per species (in total 327 plant species) expressed as cover percentage estimation according to van der Maarel (vdM 1979; 7 classes: 1-7); nomenclature according to Lauber & Wagner (1996)

en: Soil properties

en:
Plot.ID	consecutive numbers for plots (in total 36 plots)
Treatment	One restoration treatment and two reference treatments. Topsoil+Propagules= Topsoil removal (upper 10-30 cm) combined with introduction of target plant species; Initial=intensively managed grasslands; Target= species-rich semi-wet to semi-dry grasslands
treat.yr	Treatment abbreviation. For restored grasslands: Treatment abbreviation + time since restoration; Initial + Target = NA
yr.trt	year when topsoil was removed; Initial + Target = NA
Age	time since restoration (sampling year 2018 - yr.trt); Initial + Target = NA
Age.class	restored sites of similar age grouped into four age classes. Y.4 = Year 3+4 (recently restored); Y.18 = Year 17+19 (mid-term); Y.24 = Year 23+25 (long-term); Y.30 = Year 27+32 (long-term); Initial + Target = NA
Cluster	clustered spatial arrangment of sites in the field (see Supporting Information Fig. S1)
Site	clustered spatial arrangment (replicates) of plots per site (see Supporting Information Fig. S1)
N.pool	Soil nitrogen pool (June 2017); corrected for bulk density and 12 cm sampling depth [kg ha-1 12cm-1]
Corg.pool	Soil organic carbon pool (June 2017); corrected for bulk density and 12 cm sampling depth [kg ha-1 12cm-1]
pH	soil pH [CaCl2]
BD	Bulk density [g cm-3]
FC	Field capacity [%]
Veg.1	principle coordinate axis 1 (vegetation)
Veg.1	principle coordinate axis 2 (vegetation)
Nem.1	principle coordinate axis 1 (nematoda)
Nem.2	principle coordinate axis 2 (nematoda)
Fun.1	principle coordinate axis 1 (fungi)
Fun.2	principle coordinate axis 2 (fungi)
Prok.1	principle coordinate axis 1 (prokaryota)
Prok.2	principle coordinate axis 2 (prokaryota)
Veg.H	Shannon diversity (vegetation)
Nem.H	Shannon diversity (nematoda)
Fun.H	Shannon diversity (fungi)
Prok.H	Shannon diversity (prokaryota)

en: Networks

en:
Treatment	Treatment groups; Y.4-18 = restored grasslands; Initial = intensively managed grasslands; Target = species-rich semi-wet to semi-dry grasslands
Correlation	non-parametric Spearman rank correlations
Vertex.1+2	Comparison between network components; N.pool = total soil nitrogen pool [kg ha-1 12 cm-1]; Corg.pool = soil organic carbon pool [kg ha-1 12 cm-1], pH = soil pH [CaCl2]; BD = bulk density [g cm-3]; FC = field capacity [%]
	PCoA 1 + 2 biotic communities [Veg = vegetation; Nem = nematoda; Fun = fungi; Prok = prokaryota]
Type	Interactions type; A.A = abiotic.abiotic; A.B = abiotic.biotic; B.B = biotic.biotic

en: Nematode data

en:
Sample.data
Plot.ID	consecutive numbers for plots (in total 36 plots)
Treatment	One restoration treatment and two reference treatments. Topsoil+Propagules= Topsoil removal (upper 10-30 cm) combined with introduction of target plant species; Initial=intensively managed grasslands; Target= species-rich semi-wet to semi-dry grasslands
treat.yr	Treatment abbreviation. For restored grasslands: Treatment abbreviation + time since restoration; Initial + Target = NA
yr.trt	year when topsoil was removed; Initial + Target = NA
Age	time since restoration (sampling year 2018 - yr.trt); Initial + Target = NA
Age.class	restored sites of similar age grouped into four age classes. Y.4 = Year 3+4 (recently restored); Y.18 = Year 17+19 (mid-term); Y.24 = Year 23+25 (long-term); Y.30 = Year 27+32 (long-term); Initial + Target = NA
Cluster	clustered spatial arrangment of sites in the field (see Supporting Information Fig. S1)
Site	clustered spatial arrangment (replicates) of plots per site (see Supporting Information Fig. S1)

Taxonomy
ASV.ID	unit number.letter code per amplicon sequence variant (in total 904)
Taxonomic rank	Phylum, Class, Order, Family, Genus, Species

Abundance
columns	Plot.ID (see Sample.data)
rows	ASV.ID (see Taxonomy)

en:
This dataset contains all data, on which the following publication below is based.

Paper Citation:

_Resch, M.C., Schütz, M., Ochoa-Hueso, R., Buchmann, N., Frey, B., Graf, U., van der Putten, W.H., Zimmermann, S., Risch, A.C. (in review). Long-term recovery of above- and belowground interactions in restored grassland after topsoil removal and seed addition. Journal of Applied Ecology_

Please cite this paper together with the citation for the datafile.

Study area and experimental design
The study was conducted in and around two nature reserves, Eigental and Altläufe der Glatt, which were located approximately 5 km apart (47°27´ to 47°29´ N, 8°37´ to 8°32´ E, 417 to 572 m a.s.l., Canton of Zurich, Switzerland; Figure S1 and S2, Table S1). Mean annual temperature and precipitation are 9.8 ± 0.6 °C and 990 ± 168 mm (Kloten climate station 1988-2018; MeteoSchweiz, 2019). TFor this study, we used a space-for-time approach based on eight restoration sites that were between 3 and 32 years old. We measured recovery and restoration success by comparing the restored grasslands with intensively managed and semi-natural grasslands. Using a space-for-time approach requires high similarities in historical properties of the site, such as soil conditions and management regimes, to assure that temporal processes are appropriately represented by spatial patterns (Walker et al., 2010). This was the case in our study. The restored sites had similar soil conditions (i.e., soil type, structure, water availability) as the targeted semi-natural grasslands, while they shared the same agricultural legacy with intensively managed grasslands, i.e., biomass harvest and fertilization (manure and/or slurry) three to five times a year as well as tillage. We randomly established three 5 m x 5 m (25-m2) plots for plant identification and three 2 m x 2 m (4-m2) subplots for soil biotic and abiotic data collection at least 2 m away from the 25-m2 plots in each restoration site. Sites of similar age were grouped into four age classes: Y.4 (3 & 4 years after restoration), Y.18 (17 & 19 years), Y.24 (23 & 25 years), and Y.30 (27 & 32 years). Six intensively managed (Initial) and six semi-natural grassland (Target) sites complemented the experimental set-up, for a total of 36 plots. All plots were sampled under similar conditions, i.e., day of the year, air temperature, soil moisture, and time since last rain event, in June/July 2017 (intensively managed and semi-natural plots) and 2018 (restored plots).

Collection of plants and selected soil biota data
Plant species cover (in %) was visually estimated in each 25-m2 plot in mid-June (Braun-Blanquet, 1964; nomenclature: Lauber & Wagner, 1996). We calculated Shannon diversity and assessed plant community structure. We included soil microbial (fungi, procaryotes) and nematodes in our study as they represent the majority of soil biotic diversity and abundance (Bardgett & van der Putten, 2014), cover various trophic levels of the soil food web (Bongers & Ferris, 1999), and play key roles in soil functioning and ecosystem processes (Bardgett & van der Putten, 2014). In particular, soil nematodes were found to be well suited belowground indicators to evaluate recovery/development after restoration (e.g. Frouz, et al. 2008; Kardol et al., 2009; Resch et al., 2019). We randomly collected ten soil cores (2.2 cm diameter x 12 cm depths; sampler from Giddings Machine Company, Windsor, USA) in the 4-m2 subplots to assess soil nematode and microbial (fungal, prokaryotic) diversities and community structures. For soil nematodes, eight of the soil cores were combined and gently homogenized, placed in coolers and stored at 4 °C and transported to the laboratory (Netherlands Institute of Ecology, NIOO, Wageningen, Netherlands) within three days after collection. Free-living nematodes were extracted from 200 g of fresh soil using Oostenbrink elutriators (Oostenbrink, 1960). After extraction, each sample was divided into three subsamples, two for molecular identification and one to determine nematode abundance (see Resch et al., 2019). For the molecular work, two subsamples were stored in 70% ethanol (final volume 10 mL each) and transported to the laboratory at the Swiss Federal Research Institute WSL (Birmensdorf, Switzerland). Each subsample was reduced to roughly 200 μL by centrifugation and removal of the supernatant. The remaining ethanol was vaporized (65 °C for 3 h). Thereafter, 180 μL ATL buffer solution (Qiagen, Hilden, Germany) was immediately added and samples were stored at 4 °C until further processing. From these samples, nematode metagenomic DNA was extracted using the DNeasy Blood & Tissue Kit (Qiagen, Hilden, Germany) according to the manufacturer`s protocol, except for the incubation step which was run at 56 °C for 4 h. PCR amplification of the V6-V8 region of the eukaryotic small-subunit (18S) was performed with 7.5 μL of genomic DNA template (ca. 1 ng/μL) in 25 μL reactions containing 5 μL PCR reaction buffer, 2.5 mM MgCL2, 0.2 mM dNTPs, 0.8 μM of each primer (NemF: Sapkota & Nicolaisen, 2015; 18Sr2b: Porazinska et al., 2009), 0.5 μL BSA, and 0.25 μL GoTaq G2 Hot Start Polymerase (Promega Corporation, Madison, USA). Amplification was using an initial DNA denaturation step of 95 °C for 2 min, followed by 35 cycles at 94 °C for 40 sec, 58 °C for 40 sec, 72 °C for 1 min, and a final elongation step at 72 °C for 10 min. Filtering, dereplication, sample inference, chimera identification, and merging of paired-end reads was implemented using the DADA2 pipeline (v.1.12; Callahan et al., 2016) to finally assign amplicon sequence variants (ASVs) as taxonomic units.
We combined and homogenized the remaining two soil cores to assess soil microbes, placed them in coolers (4 °C) and transported them to the laboratory at WSL. Metagenomic DNA was extracted from 8 g sieved soil (2 mm) using the DNAeasy PowerMax Soil Kit (Qiagen, Hilden, Germany) according to the manufacturer´s protocol. PCR amplification of the V3-V4 region of the small-subunit (16S) of prokaryotes (i.e., bacteria and archaea) and the ribosomal internal transcribed spacer region (ITS2) of fungi was performed with 1 ng of template DNA using PCR primers and conditions as previously described (Frey et al., 2016). PCRs were run in triplicates, pooled and sent to the Genome Quebec Innovation Centre (Montreal, QC, Canada) for barcoding using the Fluidigm Access Array technology (Fluidigm) and paired-end sequencing on the Illumina MiSeq v3 platform (Illumina Inc., San Diego, USA). Quality filtering, clustering into operational taxonomic units (OTUs, 97% similarity cutoffs) and taxonomic assignment were performed as previously described (Resch et al., 2021).Taxonomic classification of nematode, prokaryotic and fungal sequences was conducted querying against the most recent versions of PR2 (v.4.11.1; Guillou et al., 2013), SILVA (v.132; Quast et al., 2013), and UNITE (v.8; Nilsson et al., 2019) reference sequence databases. Taxonomic assignment cutoffs were set to confidence rankings ≥ 0.8 (below ranked as unclassified). Prokaryotic OTUs assigned to mitochondria or chloroplasts as well as OTUs or ASVs assigned to other than Fungi or Nematoda were manually removed prior to data analysis. The three datasets were filtered to discard singletons and doubletons. Taxonomic abundance matrices were rarefied to the lowest number of sequences per community to achieve parity of the total number of reads between samples (Prokaryotes: 10,929 reads; Fungi: 18,337 reads; Nematodes: 6,662 reads). We calculated Shannon diversity and assessed community structures for soil nematodes, prokaryotes and fungi based on their relative abundances of ASV or OTU at the taxon level.

Collection of soil physical and chemical properties
We randomly collected one undisturbed soil core (5 cm diameter, 12 cm depth) per 4-m2 subplot using a steel cylinder that fit into the soil corer. The cylinders were capped to avoid disturbance during transport and used to measure field capacity, rock content and fine earth density as previously described (Resch et al., 2021). We randomly collected another three soil cores (5 cm diameter, 12 cm depths) in each 4-m2 subplot to determine soil chemical properties. The cores were pooled, dried at 60 °C for 48 h and passed through a 2 mm sieve. We measured soil pH (CaCl2) on dried samples, total nitrogen (N) and organic carbon (C) concentration on dried and fine-ground samples (≤ 0.5 mm; for details see Resch et al., 2021). We calculated total N and organic C pools after correcting its concentration for soil depth, rock content and fine earth density.

bacteria

display_name: bacteria

biodiversity

display_name: biodiversity

fungi

display_name: fungi

nematodes

display_name: nematodes

plants

display_name: plants

restoration

display_name: restoration

soil

display_name: soil

soil-biodiversity

display_name: soil-biodiversity

soil-biology

display_name: soil-biology

soil-chemistry

display_name: soil-chemistry

topsoil-removal

display_name: topsoil-removal

education

fr: Education, science

de: Bildung, Wissenschaft

en: Education and science

it: Formazione e scienza

envidat

fr: EnviDat: le portail de données environnementales

de: EnviDat: das Umwelt-Datenportal

en: EnviDat: the Environmental Data Portal

it: EnviDat: il portale dei dati ambientali

fr: EnviDat est le portail de données du WSL; il offre un accès uniformisé et réglementé aux données de surveillance et de recherche en matière d’environnement. Ce portail est capable d’héberger et de publier des ensembles de données précieuses et uniques, qui sont partagées aux quatre coins de la planète. Ce faisant, EnviDat favorise la recherche et contribue à l’avancement des sciences environnementales.

en: EnviDat is the WSL data portal providing unified and managed access to environmental monitoring and research data. The portal has the capability to host and publish data sets. Through the sharing of unique and high-value data sets worldwide, EnviDat fosters research and contributes to advancing environmental science.

de: EnviDat ist das Datenportal der WSL für einen einheitlichen und verwalteten Zugang zu Umweltdaten aus Monitoring- und Forschungsprojekten. Auf dem Portal können Datensätze gehostet und veröffentlicht werden. Durch die weltweite Bereitstellung einzigartiger und wertvoller Datensätze leistet EnviDat einen Beitrag zur Forschung im Bereich der Umweltwissenschaften.

it: EnviDat è il portale dati del WSL che fornisce un accesso unificato e gestito ai dati di monitoraggio ambientale e di ricerca. Il portale ha la capacità di ospitare e pubblicare serie di dati preziosi e unici che vengono condivisi in tutto il mondo. Così facendo, EnviDat promuove la ricerca e contribuisce al progresso della scienza ambientale.

political_level: other

wsl

capacity: public

type: organization

approval_status: approved

key: guid

key: uri

value: Envidat Harvester

en: Long-term recovery of above-and belowground interactions in restored grasslands


en: wind-speed precipitation relative-humidity long-term swiss-forest-lab meteo climate-change temperature par meteorology

en: Site aerial photo (Vordemwald)

media_type: image/jpeg

en: Site aerial photo from both Vordemwald stations. Source: Google. 2016. Site aerial photo from both Vordemwald stations. Retrieved November 11, 2016, from https://www.google.ch/maps

format: SERVICE

mimetype: image/jpeg

en: Field station data (VOF)

media_type: text/csv

en: Data file.

format: CSV

mimetype: text/csv

en: Data plot of the field station (VOF)

media_type: application/pdf

en: Data overview for air temperature, relative humidity, precipitation, PAR, wind speed.

format: PDF

mimetype: application/pdf

en: Data plot of the forest station (VOB)

en: Site photo of the forest station (VOB)

en: Site photo of the forest station from the side.

en: Forest station data (VOB)

en: Site photo of the field station (VOF)

en: Site photo of the field stationfrom the side.

en: High quality meteorological data are needed for long-term forest ecosystem research, particularly in the light of global change. The long-term data series published here comprises almost 20 years of measurements for two meteorological stations in Vordemwald in Switzerland where one station is located within a natural mixed forest stand (VOB) with European silver fir (_Abies alba_; 110 yrs) and oak trees (_Quercus sp._; 190-210 yrs) as dominant tree species. A second station is situated in the very vicinity outside of the forest (field station, VOF). The meteorological time series are presented in hourly time resolution of air temperature, relative humidity, precipitation, photosynthetically active radiation (PAR) and wind speed. Vordemwald is part of the Long-term Forest Ecosystem Research Programme (LWF) established and maintained by the Swiss Federal Research Institute WSL.

climate-change

display_name: climate-change

long-term

display_name: long-term

meteo

display_name: meteo

meteorology

display_name: meteorology

par

display_name: par

precipitation

display_name: precipitation

relative-humidity

display_name: relative-humidity

swiss-forest-lab

display_name: swiss-forest-lab

temperature

display_name: temperature

wind-speed

display_name: wind-speed

en: Vordemwald, Switzerland: Long-term forest meteorological data from the Long-term Forest Ecosystem Research Programme (LWF), from 1996 onwards


en: geochemical-solutes stable-water-isotopes spatially-distributed-sampling precipitation stream-water

en: The dataset contains daily measurements of volumes, isotopes and solutes of stream water as well as precipitation measured several locations; meteorological variables are added. Date: 18.07.2019

en:
This dataset contain measurements of solute and stable water isotopes in stream water and precipitation in the Alp catchment and two of its tributaries (between 2015 -2018) . The river Alp is a snow-dominated catchment situated in Central Switzerland characterized by an elevation range from 840 to 1898 m a.s.l. The dataset provides solutes (major anions and cations, trace metals) and stable water isotopes and water fluxes (precipitation rates, discharge) at daily intervals from several sampling locations.
An updated version of the isotope dataset is available here: https://www.doi.org/10.16904/envidat.242

geochemical-solutes

display_name: geochemical-solutes

spatially-distributed-sampling

display_name: spatially-distributed-sampling

stable-water-isotopes

display_name: stable-water-isotopes

stream-water

display_name: stream-water

education

en: Education and science

envidat

en: EnviDat: the Environmental Data Portal

en: EnviDat is the WSL data portal providing unified and managed access to environmental monitoring and research data. The portal has the capability to host and publish data sets. Through the sharing of unique and high-value data sets worldwide, EnviDat fosters research and contributes to advancing environmental science.

en: Daily data of solute and stable water isotopes in stream water and precipitation in the Alp catchment, Central Switzerland


en: literature-review stillberg treeline-ecosystems long-term-experiment mountain-ecology

media_type: .zip

en: The Stillberg bibliography (Stillberg_bibliography_v1.zip) comprises a comprehensive list of 276 scientific publications, 91 of them published in peer-reviewed ISI journals (Stillberg_bibliography_data_v1.csv) along with the metadata file (Stillberg_bibliography_metadata_v1.csv). Currently the bibliography comprises literature about the main afforestation experiment, the FACE × warming experiment, the nutrient addition experiment, and the G-TREE experiment, as well as further publications related to the Stillberg research site that have been published until August 2023. The bibliography can be filtered for different categories, e.g., experiment, peer-review, source repository or database, and source title.

en:
 Background information

The Stillberg ecological treeline research site in the Swiss Alps was established in 1975, with the aim to develop ecologically, technically, and economically sustainable reforestation techniques at the treeline to reduce the risk of snow avalanches. In the course of time, additional research aspects gained importance, such as the ecology of the treeline ecotone under global change. Over almost fifty years, research at the Stillberg site combined long-term monitoring of the large-scale high-elevation afforestation with experimental manipulations simulating global change impacts. Besides providing a scientific basis and practical guidelines for high-elevation afforestation, this research has contributed to a comprehensive understanding of ecological processes in the treeline ecotone across different compartments and scales, from individual trees, non-tree vegetation and soils to whole ecosystems, in the context of global change resulting in more than 150 publications.

 Dataset generation

We compiled a comprehensive list of scientific publications covering research at the Stillberg research site by conducting searches in the literature databases Web of Science and Google Scholar, as well as in the Digital Object Repository of the Swiss Federal Institute for Forest, Snow and Landscape Research WSL (DORA). We compiled all publications about the afforestation experiment, the FACE × warming experiment, the nutrient addition experiment, the G-TREE experiment, as well as other studies related to the Stillberg research site.

 Data description

The Stillberg bibliography (Stillberg_bibliography_data_v1.csv) comprises a comprehensive list of 276 scientific publications, 91 of them published in peer-reviewed ISI journals. Currently the bibliography comprises literature about the main afforestation experiment, the FACE × warming experiment, the nutrient addition experiment, and the G-TREE experiment, as well as further publications related to the Stillberg research site that have been published until August 2023. The bibliography can be filtered for different categories, e.g., experiment, peer-review, source repository or database, and source title. The bibliography is described in a metadata file (Stillberg_bibliography_metadata_v1.csv). The bibliography along with the metadata file are provided in a ZIP-folder (Stillberg_bibliography_v1.zip).

literature-review

display_name: literature-review

long-term-experiment

display_name: long-term-experiment

mountain-ecology

display_name: mountain-ecology

stillberg

display_name: stillberg

treeline-ecosystems

display_name: treeline-ecosystems

en: Bibliography of the long-term treeline research site Stillberg, Switzerland


maintainer: James W. Kirchner

en: streamflow precipitation stable-water-isotopes deuterium chloride catchment plynlimon tracer

en: Data Set Documentation

en: This file contains a documentation of the data sets, including references to other helpful documentations.

en: 7-hourly stable water istopes

media_type: text/plain

en: Data set of 7-hourly stable water isotope data measured at Carreg Wen (precipitation) and Upper Hafren (streamflow).

format: TXT

mimetype: text/plain

en: Weekly stable water isotopes

en: Data set of weekly stable water isotope data collected at Carreg Wen (precipitation) and Lower Hafren and Tanllwyth (streamflow).

en: The data base contains timeseries of stable water isotopes in precipitation and streamflow at Plynlimon, Wales, UK. One data set contains weekly stable water isotope data from the Lower Hafren and Tanllwyth catchments, and the other data set contains 7-hourly stable water isotope data from Upper Hafren. Both data sets also include chloride concentrations in precipitation and streamflow.

catchment

display_name: catchment

chloride

display_name: chloride

deuterium

display_name: deuterium

plynlimon

display_name: plynlimon

streamflow

display_name: streamflow

tracer

display_name: tracer

en: Stable water isotopes in precipitation and streamflow at Plynlimon, Wales, UK


en: relative-humidity climate-change swiss-forest-lab meteorology temperature precipitation wind-speed meteo par long-term

en: Site photo of the forest station (LAB)

en: Site photo of the fforest station from the side.

en: Site aerial photo (Lausanne)

en: Site aerial photo from both Lausanne stations. Source: Google. 2016. Site aerial photo from both Lausanne stations. Retrieved November 11, 2016, from https://www.google.ch/maps

en: Data plot of the field station (LAF)

en: Data overview for air temperature, relative humidity, precipitation, PAR, wind speed.

en: Data plot of the forest station (LAB)

en: Site photo of the field station (LAF)

en: Site photo of the field station from the side.

en: Forest station data (LAB)

en: Data file.

en: Field station data (LAF)

en: High quality meteorological data are needed for long-term forest ecosystem research, particularly in the light of global change. The long-term data series published here comprises almost 20 years of measurements for two meteorological stations in Lausanne in Switzerland where one station is located within a natural mixed forest stand (LAB) with European beech (_Fagus sylvatica_; 160-170 yrs), European silver fir (_Abies alba_; 160-170 yrs) and Norway spruce (_Picea abies_; 160-170 yrs) as dominant tree species. A second station is situated in the very vicinity outside of the forest (field station, LAF). The meteorological time series are presented in hourly time resolution of air temperature, relative humidity, precipitation, photosynthetically active radiation (PAR) and wind speed. Lausanne is part of the Long-term Forest Ecosystem Research Programme (LWF) established and maintained by the Swiss Federal Research Institute WSL.

climate-change

display_name: climate-change

long-term

display_name: long-term

meteo

display_name: meteo

meteorology

display_name: meteorology

par

display_name: par

precipitation

display_name: precipitation

relative-humidity

display_name: relative-humidity

swiss-forest-lab

display_name: swiss-forest-lab

temperature

display_name: temperature

education

en: Education and science

en: Lausanne, Switzerland: Long-term forest meteorological data from the Long-term Forest Ecosystem Research Programme (LWF), from 1996 onwards


maintainer: Mountain Hydrology and Mass Movements

en: debris-flow-fatalities ice-avalanche-fatalities natural-hazard-fatalities landslide-fatalities earthquake-fatalities rockfall-fatalities lightning-fatalities windstorm-fatalities flood-fatalities

media_type: CSV

en: Natural hazard fatalities per year in Swizterland since 1946.

en:
This dataset contains the number of fatalities due to flood, debris flow, landslide, rockfall, windstorm, lightning, ice avalanche, earthquake and other processes like roof avalanche or lacustrine tsunami for each year since 1946.
The following information is contained (by column and column title):

* year
* total number of hazard fatalities
* number of fatalities by flood (German: Hochwasser, Überschwemmung). Flood includes people drowned in flooded or inundated areas or carried away in streams under high-water conditions.
* number of fatalities by debris flow (German: Murgang).
* number of fatalities by landslide (German: Erdrutsch). Landslide includes people killed by landslides and hillslope debris flows (German: Hangmure).
* number of fatalities by rockfall (German: Steinschlag, Fels- und Bergsturz).
* number of fatalities by windstorm (German: Sturm). Windstorm includes people killed by falling objects or trees during very strong wind conditions and people who drowned in lakes because their boat capsized during such conditions.
* number of fatalities by lightning (German: Blitz).
* number of fatalities by ice avalanche (German: Eislawine).
* number of fatalities by earthquake (German: Erdbeben).
* number of fatalities by other processes like roof avalanche, lacustrine tsunami (German: andere Prozesse wie Dachlawine, Tsunami im See).

The data was collected based on newspaper research. For more information please refer to

_Badoux, A., Andres, N., Techel, F., and Hegg, C.: Natural hazard fatalities in Switzerland from 1946 to 2015, Nat. Hazards Earth Syst. Sci., 16, 2747-2768, https://doi.org/10.5194/nhess-16-2747-2016, 2016._

The data collection is financed by the FOEN (with exception of the collection of the avalanche fatalities). The data contains the official statistics of the FOEN on fatalities due to flood, debris flow, landslide, rock fall and avalanche.

Restrictions: The data set is not complete. Only fatalities in or around settlements and on open transportation routes are included. More precisely, fatalities were not collected, when persons exposed themselves to a great danger on purpose. Or fatalities during leisure activities which are connected to a higher risk were not included (this includes e.g. canoeing or river surfing during flood, canyoning, mountaineering, climbing, walking or driving on a closed road).

Fatalities by avalanches are collected at the WSL Institute for Snow and Avalanche Research SLF. You can download the avalanche fatalities per hydrological year [here](https://www.envidat.ch/dataset/avalanche-fatalities-switzerland-1936) and per calendar year [here](https://www.envidat.ch/dataset/avalanche-fatalities-per-calendar-year-since-1936). For a direct comparison with the fatalities presented here, please download the data set with the calendar years and do not consider fatalities in the backcountry (tour) or in terrain close to ski areas (offpiste).

debris-flow-fatalities

display_name: debris-flow-fatalities

earthquake-fatalities

display_name: earthquake-fatalities

flood-fatalities

display_name: flood-fatalities

ice-avalanche-fatalities

display_name: ice-avalanche-fatalities

landslide-fatalities

display_name: landslide-fatalities

lightning-fatalities

display_name: lightning-fatalities

natural-hazard-fatalities

display_name: natural-hazard-fatalities

rockfall-fatalities

display_name: rockfall-fatalities

windstorm-fatalities

display_name: windstorm-fatalities

envidat

en: EnviDat: the Environmental Data Portal

en: EnviDat is the WSL data portal providing unified and managed access to environmental monitoring and research data. The portal has the capability to host and publish data sets. Through the sharing of unique and high-value data sets worldwide, EnviDat fosters research and contributes to advancing environmental science.

en: Number of natural hazard fatalities per year in Switzerland since 1946


maintainer: Henning Löwe

en: microstructure discrete-element-modelling snow-mechanics unconfined-compression-experiments spherical-model-snow 3d-ct-images

en: Data Structure (Python Notebook - view online)

media_type: ipynb

en: Data Access

media_type: HTTPS

en: Data Structure (Python Notebook)

en:
For the investigation of microstructural and mechanical properties of snow unconfined compression experiments and 3D computed tomography (CT) imaging were performed on sintered rounded grain snow and spherical model snow. The spherical model snow was generated to create geometrically simplified, well-defined microstructures for calibration of numerical models, such as discrete element models (DEM) in which the microstructure is represented by spherical particles.
In the experiments, microstructural variation was created by varying the sintering time (contact size) and the density of the ice sphere samples (number of contacts). The 3D CT images allow for a complete reconstruction of the entire experimental sample (cylindrical sample dimension: diameter = 33.6 mm; height = 14 mm).

discrete-element-modelling

display_name: discrete-element-modelling

microstructure

display_name: microstructure

snow-mechanics

display_name: snow-mechanics

spherical-model-snow

display_name: spherical-model-snow

unconfined-compression-experiments

display_name: unconfined-compression-experiments

en: Unconfined compression experiments and 3D CT images of spherical model snow and RG snow samples


en: cryowrf antarctica landuse wrf topography

en: Documentation

en: Documentation about how the provided dataset was produced including details to reproduce the dataset.

en: Scripts

media_type: application/zip

en: Scripts to run the preprocessing to produce WRF binary tiles from REMA and AntarcticaLC2000 for topography and landuse (see Documentation for procedure).

format: ZIP

mimetype: application/zip

en: WPSinput

en: Input files to run the Weather Research and Forecasting model (WRF) preprocessing system WPS with REMA topography and AntarcticaLC2000 landuse data (see Documentation for details).

en:
Reference Elevation Model of Antarctica (REMA) topography and AntarcticaLC2000 landuse data
are now available as static data input for the Weather Research and Forecasting model (WRF). Topography
and landuse are made available at a spatial resolution of 1 km. This documentation describes the methods
applied to convert REMA and AntarcticaLC2000 to WRF readable format and shows how this improves
the representation of the Antarctic topography and landuse categories over coastal Antarctic regions.

antarctica

display_name: antarctica

cryowrf

display_name: cryowrf

landuse

display_name: landuse

topography

display_name: topography

wrf

display_name: wrf

en: REMA topography and AntarcticaLC2000 for WRF


irkis-soil-moisture-measurements-davos

maintainer: Nander Wever

title_for_slug: irkis-soil-moisture-measurements-davos

en: meteo-station dischma-catchment soil-moisture davos meteorological-measurements

en: IRKIS soil moisture data (zip file)

en: Compressed archive (zip) containing soil moisture measurements, as well as meteorological measurements and interpolated meteorological time series.

en:
Meteorological and soil moisture measurements from soil moisture stations installed from October 2010 - October 2013 in the area surrounding Davos, in particular in the Dischma catchment.

There are in total 7 stations: 1202, 1203, 1204, 1205, 222, 333 and SLF2. For each of the stations, there is a:

* vwc_[stn].smet: containing the soil moisture measurements

* station_[stn].smet: in-situ measured meteorlogical parameters. Note, the quality of these measurements for stations 1202, 1203, 1204 and 1205 is very low, with data gaps. Use this data with care. For stations 222, 333 and SLF2, data quality is high and only the default cautiousness should be applied.

* interpolatedmeteo_[stn].smet contains per stations a dataset derived by interpolating from several stations in the Davos area to the stations location. This dataset was generated from the output of the Alpine3D model, of which simulations are presented in the Wever et al. (2017) manuscript.

At the soil moisture measurement sites, Decagon 10HS sensors were installed, at 10, 30, 50, 80 and 120 cm depth. Per depth 2 sensors were installed, labelled A and B in the datafiles. Note that at stations 1203, 1204 and 1205, sensors were only installed at 10, 30 and 50 cm depth.

The files follow the SMET format: https://models.slf.ch/docserver/meteoio/SMET_specifications.pdf and metadata for the stations can be found in the header of the smet files.

Please cite the Wever et al. (2017) reference when using this data in publications.

For a more detailed description, please refer to: Wever, N., Comola, F., Bavay, M., and Lehning, M.: Simulating the influence of snow surface processes on soil moisture dynamics and streamflow generation in an alpine catchment, Hydrol. Earth Syst. Sci., 21, 4053-4071, https://doi.org/10.5194/hess-21-4053-2017, 2017.

davos

display_name: davos

dischma-catchment

display_name: dischma-catchment

meteo-station

display_name: meteo-station

meteorological-measurements

display_name: meteorological-measurements

soil-moisture

display_name: soil-moisture

education

en: Education and science

envidat

en: EnviDat: the Environmental Data Portal

en: EnviDat is the WSL data portal providing unified and managed access to environmental monitoring and research data. The portal has the capability to host and publish data sets. Through the sharing of unique and high-value data sets worldwide, EnviDat fosters research and contributes to advancing environmental science.

en: IRKIS Soil moisture measurements Davos


ceilometer-klosters

title_for_slug: ceilometer-klosters

en: vertical-visibility klosters raclets ceilometer cloud-base-height

en: Ceilometer data Klosters

media_type: mat

en: Ceilometer installed at Klosters measuring cloud base height, cloud cover and backscatter profile. The time resolution is 30 seconds, with time given in number of days from January 0, 0000, UTC. More information on the variables in the manufacturer manual.

en: Cloud base height (m) and vertical visibility (m) were measured with the VAISALA Ceilometer CL31 in Klosters (LON: 9.880413, LAT: 46.869019). The CL31 is an instrument with constant reliability for all weather conditions and simultaneous detection of three cloud layers in heights up to 7.6 km.

ceilometer

display_name: ceilometer

cloud-base-height

display_name: cloud-base-height

klosters

display_name: klosters

raclets

display_name: raclets

vertical-visibility

display_name: vertical-visibility

en: Ceilometer Klosters


photogrammetric-drone-data-schurlialp

title_for_slug: photogrammetric-drone-data-schurlialp

en: uav wingtra orthophoto photogrammetry drone-data dsm

en: Data Access

media_type: TIF

en:
- 10 cm DSM
- 10 cm HS
- 3 cm orthophoto
- original RGB images
- Agisoft Metashape Processing Report

All of the products are also available as a cloud optimized geotiff.

en: The data was collected on 16.04.2021 and on 28.05.2021 with a Wingtra Gen II and a Sony RX1 II RGB sensor to obtain snow depth and distribution data. Following the data collection, the data was processed with Agisoft Metashape. A 10cm DSM, a 10cm snow depth raster, a 3mm orthophoto and the original drone images are available for download.

drone-data

display_name: drone-data

dsm

display_name: dsm

orthophoto

display_name: orthophoto

photogrammetry

display_name: photogrammetry

uav

display_name: uav

wingtra

display_name: wingtra

en: Photogrammetric Drone Data Schürlialp


en: outgoing shortwave-radiation longwave-radiation davos incoming

en: Processing doc

en: DATA ACCESS

media_type: SMET

en: Parameter file

en: Metadata

en:
Incoming and outgoing shortwave and longwave 2 min radiation measurements in Davos Dorf, CH.

 References
1. Marty, C., Philipona, R., Frohlich, C., Ohmura, A.. Altitude dependence of surface radiation fluxes and cloud forcing in the alps: results from the alpine surface radiation budget network. 2002. Theoretical and Applied Climatology. Volume 72. Issue 3-4. 137-155. http://dx.doi.org/10.1007/s007040200019. 10.1007/s007040200019.
2. Christoph Marty. Surface Radiation, Cloud Forcing and Greenhouse Effect in the Alps. 2000. Institute fuer Klimaforschung ETH. Zuercher Klima-Schriften. Volume 79. http://e-collection.library.ethz.ch/eserv/eth:23491/eth-23491-01.pdf.

incoming

display_name: incoming

longwave-radiation

display_name: longwave-radiation

outgoing

display_name: outgoing

shortwave-radiation

display_name: shortwave-radiation

en: ASRB_DAV: Shortwave and longwave radiation measurements (2 min) in Davos Dorf


en: temperature swiss-forest-lab climate-change relative-humidity wind-speed long-term par meteo precipitation meteorology

en: Site photo of the forest station (LEB)

en: Site photo of the forest station from below (viewing upwards the slope)

en: Forest station data (LEB)

en: Data file.

en: Data plot of the forest station (LEB)

en: Data overview for air temperature, relative humidity, precipitation, PAR, wind speed.

en: High quality meteorological data are needed for long-term forest ecosystem research, particularly in the light of global change. The long-term data series published here comprises almost 20 years of measurements for one meteorological station in Lens in Switzerland which is located within a natural coniferous forest with Scots pine (_Pinus sylvestris_; 150-170 yrs)) as dominant tree species. The meteorological time series are presented in hourly time resolution of air temperature, relative humidity, precipitation, photosynthetically active radiation (PAR) and wind speed. Lens is part of the Long-term Forest Ecosystem Research Programme (LWF) established and maintained by the Swiss Federal Research Institute WSL.

climate-change

display_name: climate-change

long-term

display_name: long-term

meteo

display_name: meteo

meteorology

display_name: meteorology

par

display_name: par

precipitation

display_name: precipitation

relative-humidity

display_name: relative-humidity

swiss-forest-lab

display_name: swiss-forest-lab

temperature

display_name: temperature

wind-speed

display_name: wind-speed

education

en: Education and science

envidat

en: EnviDat: the Environmental Data Portal

en: EnviDat is the WSL data portal providing unified and managed access to environmental monitoring and research data. The portal has the capability to host and publish data sets. Through the sharing of unique and high-value data sets worldwide, EnviDat fosters research and contributes to advancing environmental science.

en: Lens, Switzerland: Long-term forest meteorological data from the Long-term Forest Ecosystem Research Programme (LWF), from 1997 onwards


en: radiation shortwave-radiation longwave-radiation davos weisfluhjoch

en: DATA ACCESS

en: ASRB radiation station at the Weissfluhjoch versuchsfeld.

en: Processing doc

en: Description of processing code that was ported to R by P. Frei (2016).

en: Description of ASRB parameters.

en:
Incoming and outgoing shortwave and longwave 2 min radiation measurements at the Weissfluhjoch research site, Davos, CH. The experimental site at the Weissfluhjoch (WFJ, 46.83 N, 9.81 E) is located at an altitude of 2540 m in the Swiss Alps near Davos. During the winter months, almost all precipitation falls as snow at this altitude. As a consequence, a continuous seasonal snow cover builds up every winter, with a maximum snow height ranging from 153–366 cm over the period 1934–2012. The measurement site is located in an almost flat part of a southeast oriented slope.

 References
1. Marty, C., Philipona, R., Frohlich, C., Ohmura, A.. Altitude dependence of surface radiation fluxes and cloud forcing in the alps: results from the alpine surface radiation budget network. 2002. Theoretical and Applied Climatology. Volume 72. Issue 3-4. 137-155. http://dx.doi.org/10.1007/s007040200019. 10.1007/s007040200019.
2. Christoph Marty. Surface Radiation, Cloud Forcing and Greenhouse Effect in the Alps. 2000. Institute fuer Klimaforschung ETH. Zuercher Klima-Schriften. Volume 79. http://e-collection.library.ethz.ch/eserv/eth:23491/eth-23491-01.pdf.

davos

display_name: davos

radiation

display_name: radiation

weisfluhjoch

display_name: weisfluhjoch

en: ASRB_WFJVF: Shortwave and longwave radiation measurements (2 min) at the Weissfluhjoch research site, Davos


en: functional-diversity ecological-indicator-values taxonomic-diversity climate grain-size soil phylogenetic-diversity

en: Data access

media_type: ZIP

en:
This dataset comprises a large array of ecological data for the European Alps:

  (1) Current soil and climate predictors at various resolutions.
  (2) GBIF observations of the European Alps Flora (~4,000 species).
  (3) Species habitat suitability maps (1,109 species; based on species observations filtered at 40x40-km)
  at various resolutions used in the study to generate (4); except 'expert'...
  (4) Expert, Taxonomic, phylogenetic and functional diversity of the study region at various resolutions
  (from 100-m to 40-km --> 100-m aggregated & mean to km + non-aggregated/predicted) for CLIM,
  SOIL and CLIM-SOIL models.
  (5) Ecological and altitudinal preferences of the European Alps Flora.
  (6) Data outputs of the related published article.
  (7) All scripts used for analyses.
  (8) Additional files used for analyses.
  (9) Improved set of species habitat suitability maps (~2,600 species; based on species observations
  filtered at 1x1-km) and related taxonomic diversity at 100-m resolution (aggregated to km +
  non-aggregated/predicted) for CLIM, SOIL and CLIM-SOIL models ---> not incorporated in the study.

climate

display_name: climate

ecological-indicator-values

display_name: ecological-indicator-values

functional-diversity

display_name: functional-diversity

grain-size

display_name: grain-size

phylogenetic-diversity

display_name: phylogenetic-diversity

soil

display_name: soil

taxonomic-diversity

display_name: taxonomic-diversity

en: Resolution in species distribution models shapes spatial patterns of plant multifaceted diversity


high-resolution-climate-data-for-europe

title_for_slug: high-resolution-climate-data-for-europe

en: air-temperature eur11 cordex maximum-temperature precipitation europe daily-climate minimum-temperature

en: CHELSA EUR11 Technical Documentation

media_type: PDF

en: As the CHELSA project is continuously expanded, certain new parameters, time scales etc. are not included anymore in the original publication (Karger et al. 2017). We are therefore provide this document as a guideline of the current climatic parameters available and will expand it as new parameters or time periods will become available. Please refer to the version history for changes made in the document.

en: Data Access

media_type: NetCDF

en:
High-resolution information on climatic conditions is essential to many applications in environmental and ecological sciences. Here we present downscaled climate data for the CORDEX EUR11 domain at a high resolution of 30 arc sec. The temperature algorithm is based on statistical downscaling of atmospheric temperature lapse rates. The precipitation algorithm incorporates orographic predictors including wind fields, valley exposition, and boundary layer height. The resulting data consist of a daily temperature and precipitation timeseries.

The data is distributed under a: Creative Commons: Attribution 4.0 International (CC BY 4.0) license.

air-temperature

display_name: air-temperature

cordex

display_name: cordex

daily-climate

display_name: daily-climate

europe

display_name: europe

maximum-temperature

display_name: maximum-temperature

minimum-temperature

display_name: minimum-temperature

en: High resolution climate data for Europe


en: dispersal biodiversity reef-fish ecology traits

en: Input for the Global distribution maps for proportions of species richness for (a) simulated dispersal parameter value (b) observed dispersal-related traits

en: Geographic distribution data (species richness and proportions of high/low dispersal-related traits) on predominantly low dispersing and high dispersing species expected based on the simulations, and those where low- and high- dispersal related traits values are observed for the five investigated traits.

en: Donati et al. main script

media_type: R

en:
Main script used in : Donati et al. 2019 "A process-based-model supports an association between dispersal and the prevalence of species traits in tropical reef fish assemblages"
accepted in Ecography, August 2019

en: Global simulated distribution of species under a range of dispersal parameter values

en:
Global distribution of simulated species richness under a range of dispersal parameter values d
d ∈ [1.75°: 5.45°].

en: Dispersal parameter value d per simulation starting with a minimum dispersal value, where each species is able to colonise each adjacent cell, and ending with the value of d for which all the extant habitats, including the most remote Pacific islands, have been colonised (d ∈ [1.75°: 5.45°]).

en: Supplementary material Appendix 1 Figures illustrated in Donati et al. 2019 "A process-based-model supports an association between dispersal and the prevalence of species traits in tropical reef fish assemblages". accepted in Ecography, August 2019

en: Main text figures

en: R Script used for figures in main text in: Donati et al. 2019: "A process-based-model supports an association between dispersal and the prevalence of species traits in tropical reef fish assemblages" accepted in Ecography, August 2019.

en: Global coast files from https://www.ngdc.noaa.gov/mgg/shorelines/gshhs.html used in the Figures

media_type: rds

en: Simulated rates of speciation and extinction under a range of dispersal parameters. Net diversification rates are calculated from the difference between speciation and extinction rates. This data allows the creation of Figure A4 in Supporting information, Appendix 1.

en: Coral reef grid cells (shape, dbf, etc. files)

en:
Grid cells of 5 × 5°, corresponding to circa 555 × 555 km at the Equator (see Parravicini et al. 2013, Leprieur et al. 2016)

Parravicini, V. et al. 2013. Global patterns and predictors of tropi- cal reef sh species richness. – Ecography 36: 1254–1262.

Leprieur, F. et al. 2016. Plate tectonics drive tropical reef biodiver- sity dynamics. – Nat. Commun. 7: 11461.

en: Tropical reef fish trait data

en:
Species trait (i.e. adult body size, home range mobility behaviour, circadian activity and schooling) data gathered from the General Approach to Species Abundance Relationships (GASPAR) database (Kulbicki et al. 2013, Parravicini et al. 2013).

Kulbicki, M. et al. 2013. Global biogeography of reef shes: a hier- archical quantitative delineation of regions. – PLoS One 8: e81847.

Parravicini, V. et al. 2013. Global patterns and predictors of tropi- cal reef sh species richness. – Ecography 36: 1254–1262.

To complement the trait data with Pelagic Larval Duration information refer to the following publication:

Luiz, Osmar J., et al. "Adult and larval traits as determinants of geographic range size among tropical reef fishes." Proceedings of the National Academy of Sciences 110.41 (2013): 16498-16502.

In Donati et al. "A process-based model supports an association between dispersal and the prevalence of species traits in tropical reef fish assemblages" accepted in Ecography in August 2019, we used the PLD gathered from Luiz et al. 2013 as described in the methods - section "Species trait data".

en: Functions used in R scripts in: Donati et al. 2019 "A process-based-model supports an association between  # dispersal and the prevalence of species traits in tropical reef fish assemblages" accepted in Ecography, August 2019

en: Functions for Figures illustrated in Donati et al. 2019 "A process-based-model supports an association between dispersal and the prevalence of species traits in tropical reef fish assemblages" accepted in Ecography, August 2019.

en: Input for spatial congruence test overlap between proportions of low and high dispersal and observed low- and high- dispersal-related traits in assemblages (30-50% threshold). see Table 2.

en: Global species distribution of tropical reef fish species

en: Each species corresponds to a "Gaspar_code" gathered from the General Approach to Species Abundance Relationships (GASPAR) database (Kulbicki et al. 2013, Parravicini et al. 2013) and is linked to a specific Fishned ID corresponding to a specific Latitude and Longitude.

en:
This dataset contains all data and R codes (R Development Core Team, https://www.R-project.org) used in the following publication: Donati GFA, Parravicini V, Leprieur F, Hagen O, Gaboriau T, Heine C, Kulbicki M, Rolland J, Salamin N, Albouy C, Pellissier L. "A process-based model supports an association between dispersal and the prevalence of species traits in tropical reef fish assemblages" accepted by Ecography in August 2019. When using this data and R scripts the above publication should be cited.

The interaction of habitat dynamics with species dispersal abilities could generate gradients
in species diversity and prevalence of life-history and ecological traits, when the latter are associated with dispersal potential. In this dataset, we use a spatial mechanistic model of speciation, extinction and dispersal, constrained by a dispersal parameter. This model allows to simulate the interplay between reef habitat dynamics over the past 140 million years and dispersal, shaping lineage diversification history and global assemblage composition of over 6000 tropical reef fish species. Global trait distribution data of tropical reef fish are used to evaluate the congruence between simulations and observations.

biodiversity

display_name: biodiversity

dispersal

display_name: dispersal

ecology

display_name: ecology

reef-fish

display_name: reef-fish

traits

display_name: traits

education

en: Education and science

envidat

en: EnviDat: the Environmental Data Portal

en: EnviDat is the WSL data portal providing unified and managed access to environmental monitoring and research data. The portal has the capability to host and publish data sets. Through the sharing of unique and high-value data sets worldwide, EnviDat fosters research and contributes to advancing environmental science.

en: Simulated and observed prevalence of dispersal-related traits in tropical reef fish assemblages worldwide


en: digital-planning wms-data cable-yarding swiss-geodata qgis-plugin geoinformation-system seilaplan open-source digital-elevation-model

en: WMS-Layer als Hintergrundkarten laden

en: View video on youtube!

en:
In order to digitally plan a cable line using the QGIS plugin ‘Seilaplan’, maps with various background information are helpful. In this tutorial we show you how to obtain maps that are helpful for cable line planning, for example a national map of Switzerland at different scales, the NFI vegetation height model or the NFI forest mix rate. For this we explain what WMS datasets are and how to integrate them into QGIS. No download of large data is needed for this, only a good internet connection. Please note that the tutorial language is German!

Link for the integration of WMS data: https://wms.geo.admin.ch/

Link to the description on the Swisstopo website:
https://www.geo.admin.ch/en/geo-services/geo-services/portrayal-services-web-mapping/web-map-services-wms.html

Link to the Seilaplan website: https://seilaplan.wsl.ch

**************************

Für die Verwendung des QGIS Plugins Seilaplan zur digitalen Seillinienplanung sind verschiedene Hintergrundkarten hilfreich. In diesem Tutorialvideo zeigen wir, was WMS Daten sind und wie man diese in QGIS einbinden kann. Dafür müssen die Daten nicht heruntergeladen werden. Es braucht lediglich eine gute Internetverbindung. Für die Seillinienplanung hilfreiche Karten sind bspw. die Landeskarte der Schweiz in verschiedenen Massstäben, das Vegetationshöhenmodell LFI oder der Waldmischungsgrad LFI.

Link zur Einbindung der WMS Daten: https://wms.geo.admin.ch/

Link zur Beschreibung auf der Swisstopo Webseite: https://www.geo.admin.ch/de/geo-dienstleistungen/geodienste/darstellungsdienste-webmapping-webgis-anwendungen/web-map-services-wms.html

Link zur Seilaplan-Website: https://seilaplan.wsl.ch

cable-yarding

display_name: cable-yarding

digital-elevation-model

display_name: digital-elevation-model

digital-planning

display_name: digital-planning

geoinformation-system

display_name: geoinformation-system

open-source

display_name: open-source

qgis-plugin

display_name: qgis-plugin

seilaplan

display_name: seilaplan

swiss-geodata

display_name: swiss-geodata

wms-data

display_name: wms-data

en: Seilaplan Tutorial: Load WMS layers as background maps


en: tartu zurich angiospermae antwerp paris floral-resources floral-inventories plants poznan urban

media_type: .csv

en: Floral units, mean number of flowers per floral unit and floral abundance of the different plants counted in quadrats in the study sites in Zurich during the four sampling periods

en: Trait values for the sampled plants

en: List of plant species sampled in the five cities (Antwerp, Paris, Poznan, Tartu and Zurich) during the different sampling perionds (four for Zurich, three for the other cities) with their frequency (oc1-oc16) within a 100 m radius

en: Data of a survey of flowering plants in 80 sites in five European cities and urban agglomerations (Antwerp, Belgium; greater Paris, France; Poznan, Poland; Tartu, Estonia; and Zurich, Switzerland) sampled between April and July 2018.

angiospermae

display_name: angiospermae

antwerp

display_name: antwerp

floral-inventories

display_name: floral-inventories

floral-resources

display_name: floral-resources

paris

display_name: paris

plants

display_name: plants

poznan

display_name: poznan

tartu

display_name: tartu

urban

display_name: urban

zurich

display_name: zurich

en: Flowering Plants (Angiospermae) in Urban Green Areas in five European Cities


maintainer: Joan Casanelles Abella

en: dna-metabarcoding zurich plant-pollinator-interactions cavity-nesting-bees osmia-bicornis trap-nest antwerp chelostoma-florisomne tartu poznan paris hylaeus-communis urban-ecology osmia-cornuta

en: Larval food composition

en: Explanation data variables

en: Urbanization poses threats and opportunities for the biodiversity of wild bees. A main gap relates to the food preferences of wild bees in urban ecosystems, which usually harbour large numbers of plant species, particularly at the larval stage. This data sets describes the larval food of four wild bee species (i.e. Chelostoma florisomne, Hylaeus communis, Osmica bicornis and Osmia cornuta) and three genera (i.e. Chelostoma sp., Hylaeus sp, and Osmia sp.) common in urban areas in five different European cities (i.e. Antwerp, Paris, Poznan, Tartu and Zurich). This data results from a European-level study aimed at understanding the effects of urbanization on biodiversity across different cities and citiscapes, and a Swiss project aimed at understanding the effects of urban ecosystems in wild bee feeding behaviour. Wild bees were sampled using standardized trap-nests in 80 sites (32 in Zurich and 12 in each of the remaining cities), selected following a double gradient of available habitat at local and landscape scales. Larval pollen was obtained from the bee nests and identified using DNA metabarconding. The data provides the plant composition at the species or genus level of the different bee nests of the studied species in the studied sites of the five European cities. For Hylaeus communis, this is the first study in reporting larval food composition.

cavity-nesting-bees

display_name: cavity-nesting-bees

chelostoma-florisomne

display_name: chelostoma-florisomne

dna-metabarcoding

display_name: dna-metabarcoding

hylaeus-communis

display_name: hylaeus-communis

osmia-bicornis

display_name: osmia-bicornis

osmia-cornuta

display_name: osmia-cornuta

plant-pollinator-interactions

display_name: plant-pollinator-interactions

trap-nest

display_name: trap-nest

urban-ecology

display_name: urban-ecology

en: Larval food composition of four wild bee species in five European cities