maintainer: Anita Risch

en: invertebrates ecosystem-coupling grassland exclusion vertebrates grazing ecosystem-functions swiss-national-park subalpine ecosystem-multifunctionality

en: Datafile

en:
ID: Plot ID composited of Fence & Treatment number (90 plots total)

Grassland: Name of the six subalpine grasslands where the exclosure setups were erected. In large grasslands (GRIM, MING, STAB), we established four hierarchical exclosure setups (two in short- and two in tall-grass vegetation), whereas in the small grasslands (BOTS, DADA, MARG) only one setup was erected per vegetation type. BOTS = Val dal Botsch, GRIM = Grimmels, MING = Val Mingér, STAB = Alp Stabelchod, DADA = Stabelchod da daint, MARG = Margunet

Fence: Number of exclosure setup (1 - 18). Note that uneven numbers (1,3,5,7,9,11,13,15,17) represent setups established on short-grass vegetation, even numbers (2,4,6,8,10,12,14,16,18) setups established in tall-grass vegetation
Vegetation	Vegetation type, 1 = short-grass vegetation, 2 = tall-grass vegetation; (9 replicate fences on each vegetation type)
Treatment	Progressive animal exclusion, numbers indicate which animals had access to the plot: 1 = Large/medium/small mammals, and invertebrates have access ("L/M/S/I"), 2 = Medium/small mammals, and invertebrates have access (“M/S/I”), 3 = Small mammals and invertebrates have access (“S/I”), 4 = Invertebrates have access (“I”), 5 = No animals have access (“None”)

Bulkden:	Fine fraction soil bulk density (g cm-3)

Soil.moist	: Soil moisture (%)

Soil.org.C: Rhizosphere soil C content (%)

pH: Soil pH

Nema.PC1: First principal component (PC1) of principal component analysis (PCA) of the soil nematode community composition

Nema.PC2: Second principal component (PC2) of principal component analysis (PCA) of the soil nematode community composition

Micro.PC1: 	First principal component (PC1) of principal component analysis (PCA) of the soil microbial community composition

Micro.PC2: Second principal component (PC2) of principal component analysis (PCA) of the soil microbial community composition

Abv.Inv.PC1: First principal component (PC1) of principal component analysis (PCA) of the aboveground-dwelling invertebrate community composition

Abv.Inv.PC2: Second principal component (PC2) of principal component analysis (PCA) of the aboveground-dwelling invertebrate community composition

Arthr.PC1: First principal component (PC1) of principal component analysis (PCA) of the soil arthropod community composition

Arthr.PC2: Second principal component (PC2) of principal component analysis (PCA) of the soil arthropod community composition

Plant.PC1: First principal component (PC1) of principal component analysis (PCA) of the plant community composition

Plant.PC2: Second principal component (PC2) of principal component analysis (PCA) of the plant community composition

Root.biomass: Root biomass (g m-2)

Micro.biomass	: Microbial biomass carbon (mg kg-1)

Plant.richness: Plant species richness

Plant.N: Plant tissue nitrogen content (%)

Soil.N.min: Soil net nitrogen mineralisation (kg ha-1 season)

Soil.resp: Soil carbon dioxide emissions (soil respiration; g m-2 h-1)

en:
This dataset contains all data on which the following publication below is based.

Paper Citation:
 > Risch AC, Ochoa-Hueso R, van der Putten WH, Bump JK, Busse MD, Frey B, Gwiazdowicz DJ, Page-Dumroese DS, Vandegehuchte ML, Zimmermann S, Schütz M. Size-dependent loss of aboveground animals differentially affects grassland ecosystem coupling and functions. 2018. Nature Communications 9: 3684. [doi: 10.1038/s41467-018-06105-4](https://doi.org/10.1038/s41467-018-06105-4).

Please cite this paper together with the citation for the datafile.

#Methods
Study sites
The experimental exclosure setups were installed within the SNP (IUCN category Ia preserve; Dudley 2008), in south-eastern Switzerland. The park covers 172 km2 of forests and subalpine and alpine grasslands along with scattered rock outcrops and scree slopes. The entire area has been protected from human impact (no hunting, fishing, camping or off-trail hiking) since 1914. Large, fairly homogenous patches of short- and tall-grass vegetation, which originate from different historical management and grazing regimes, cover the park’s subalpine grasslands entirely. Short-grass vegetation developed in areas where cattle used to rest (nutrient input) prior to the park’s foundation (14th century to 1914) (Schütz and others 2003, 2006) and is dominated by lawn grass species such as Festuca rubra L., Briza media L. and Agrostis capillaris L. (Schütz and others 2003, 2006). Today, this vegetation type is intensively grazed by diverse vertebrate and invertebrate communities that inhabit the park and consume up to 60% of the available biomass (Risch and others 2013). Tall-grass vegetation developed where cattle formerly grazed, but did not rest, and is dominated by rather nutrient-poor tussocks of Carex sempervirens Vill. and Nardus stricta L. (Schütz and others 2003, 2006). This vegetation type receives considerably less grazing, with only roughly 20% of the biomass consumed (Risch and others 2013). Consequently, the two vegetation types together represent a long-term trajectory of changes in grazing regimes. Underlying bedrock of all grasslands is dolomite, which renders these grasslands rather poor in nutrients regardless of former and current land-use regimes.
Experimental design
To progressively exclude aboveground vertebrate and invertebrate animals, we established 18 size-selective exclosure setups (nine in short-grass, nine in tall-grass vegetation) distributed over six subalpine grasslands across the SNP (Risch and others 2013, 2015). Elevation differences of exclosure locations did not exceed 350 m (between 1975 and 2300 m a.s.l.). The exclosures were established immediately after snowmelt in spring 2009 and were left in place for five consecutive growing seasons (until end of 2013). They were, however, temporarily dismantled every fall (late October after first snowfall) to protect them from avalanches. They were re-established in the same location every spring immediately after snowmelt. Each size-selective exclosure setup consisted of five plots (2 x 3 m) that progressively excluded aboveground vertebrates and invertebrates from large to small. The plots are labelled according to the guilds that had access to them “L/M/S/I”, “M/S/I”, “S/I”, “I”, “None”; L = large mammals, M = medium mammals, S = small mammals, I = invertebrates, None = no animals had access. As we only had permission to have the experimental setup in place for five consecutive growing seasons, the experiment had to be completely dismantled in the late fall of 2013 and all material removed from the SNP.
Our exclosure design was aimed at excluding mammalian herbivores, but naturally also excluded the few medium and small mammalian predators, as well as the entire aboveground invertebrate food web. A total of 26 large to small mammal species can be found in the SNP, but large apex predators are missing (wolf, bear, lynx) . Reptiles, amphibians and birds are scarce to absent in the subalpine grasslands under study. Only two reptile species occur in the park and they are confined to rocky areas that warm up enough for them to survive. One frog species spawns in an isolated pond far from our grasslands. Only three bird species occasionally feed on the subalpine grasslands. Using game cameras (Moultrie 6MP Game Spy I-60 Infrared Digital Game Camera, Moultrie Feeders, Alabaster, AL, USA), we did observe that the medium- and small-sized mammals (marmot/hares and mice) were not afraid to enter the fences and feed on their designated plots. We never spotted reptiles, amphibians or birds on camera. We distinguished between 59 higher aboveground-dwelling invertebrate taxa that our size-selective exclosures excluded (see also methods for aboveground-dwelling invertebrates below).
The “L/M/S/I” plot (not fenced) was located at least 5 m from the 2.1 m tall and 7 x 9 m large main electrical fence that enclosed the other four plots. The bottom wire of this fence was mounted at 0.5 m height and was not electrified to enable safe access for medium and small mammals, while fencing out the large ones. Within each main fence, we randomly established four 2 x 3 m plots separated by 1-m wide walkways from one another and from the main fence line: 1) the “M/S/I” plots were unfenced, allowing access to all but the large mammals; 2) the “S/I” plots (10 x 10 cm electrical mesh fence) excluded all medium-sized mammals. Note that the bottom 10 cm of this fence remained non-electrified to enable safe access for small mammals; 3) the “I” plots (2 x 2 cm metal mesh fence) excluded all mammals. We double-folded the mesh at the bottom 50 cm to reduce the mesh size to smaller than 1 x 1 cm openings; and 4) the “None” plots were surrounded by a 1 m tall mosquito net (1.5 x 2 mm) to exclude all animals. The top of the plot was covered with a mosquito-meshed wooden frame mounted to the corner posts (roof). We treated these plots a few times with biocompatible insecticide (Clean kill original, Eco Belle GmbH, Waldshut-Tiengen, Germany) to remove insects that might have entered during data collection or that hatched from the soil, but amounts were negligible and did not impact soil moisture conditions within these plots.
To assess whether the design of the “None” exclosure (mesh and roof) affected the response variables within the plots and, therefore, influenced the results, we established an additional six “micro-climate control” exclosures (one in each of the six grasslands) (Risch and others 2013, 2015). These exclosures were built as the “None” exclosures but were open at the bottom (20 cm) of the 3-m side of the fence facing away from the prevailing wind direction to allow invertebrates to enter. A 20-cm high and 3-m long strip of metal mesh was used to block access to small mammals. Thus, this construction allowed a comparable micro-climate to the “None” plots, but also a comparable feeding pressure by invertebrates to the “I” plots. We compared various properties within these exclosures against one another to assess if our construction altered the conditions in the “None” plots. We showed that differences in plant (e.g., vegetation height, aboveground biomass) and soil properties (e.g., soil temperature, moisture) found between the “I” and the “None” treatments were not due to the construction of the “None” exclosure, but a function of animal exclusions, although the amount of UV light reaching the plant canopy was significantly reduced (Risch and others 2013).
Aboveground invertebrate sampling
Aboveground invertebrates were sampled with two different methods to capture both ground- and plant-dwelling organisms: 1) we randomly placed two pitfall traps (67 mm in diameter, covered with a roof) filled with 20% propylene glycol in one 1 x 1 m subplot of the 2 x 3 m treatment plots in spring 2013 (May) and emptied them every two weeks until late September 2013 (Vandegehuchte and others 2017b, 2018). A pitfall trap consisted of a plastic cylinder (13 cm depth, 6.75 cm diameter). Within each cylinder we placed a 100 ml plastic vial with outer diameter 6.70 cm and on top of the cylinder we placed a plastic funnel to guide the invertebrates into the vials. Each trap was cover with a cone-shaped and transparent plastic roof to protect the trap from rain (Vandegehuchte and others 2017b, 2018). Note that in the “None” plots only one trap was placed as control to check for effectiveness of the exclosure. 2) We vacuumed all invertebrates from a 60 x 60 cm area on another 1 x 1 m subplot with a suction sampler (Vortis, Burkhard manufacturing CO, Ltd., Rickmansworth, Hertfordshire, UK) every month from June to September 2013 (Vandegehuchte and others 2017b, 2018). For this purpose, we quickly placed a square plastic frame (60 x 60 x 40 cm) with a closable mosquito mesh sleeve attached to the top edge into the plot from the outside. The suction sample was then inserted into through the sleeve and operated for 45 s to collect the invertebrates (Vandegehuchte and others 2017b, 2018).
We sorted the ≈100 000 individuals collected with both methods by hand and identified each individual morphologically to the lowest taxonomic level feasible (59 taxa, including orders, suborders, subfamilies, families; phylum for Mollusca). These taxa belonged to the following feeding types: 19 herbivores, 16 detritivores, 9 predators, 8 mixed feeders, 5 omnivores and 2 non-classified feeders (or not feeding as adults) (Vandegehuchte and others 2017b). We summed the numbers from the two pitfall traps and the suction sampling over the course of the 2013 season to represent the aboveground invertebrate abundance and community composition of a plot. Note: we did not specifically attempt to catch flying invertebrates with e.g., sticky traps, thus a few flying insects may have been missed with our vacuum sampling approach.
Sampling of plant properties
The vascular plant species composition was assessed at peak biomass every summer (July) by estimating the frequency of occurrence of each species with the pin count method in each plot (Frank and McNaughton 1990). A total of 172 taxa occurred within our 90 plots and we calculated plant species richness for each plot separately. We used the 2013 data in this study. Plant quality was assessed every year in July and September; here we use plant quality at the end of the experiment (September 2013). Two 10 x 100 cm wide strips of vegetation per plot were clipped, combined, dried at 65°C, and ground (Pulverisette 16, Fritsch, Idar-Oberstein, Germany) to pass through a 0.5 mm sieve. Twenty randomly selected samples across all treatments were analysed for N (Leco TruSpec Analyser, Leco, St. Joseph, Michigan, USA) (Vandegehuchte and others 2015). Nitrogen concentrations of the other samples were then estimated from models established for the experiment and the entire SNP relating Fourier transform-near infrared reflectance (FT-NIR) spectra to the measured values of N using a multi-purpose FT-NIR spectrometer (Bruker Optics, Fällanden, Switzerland) (Vandegehuchte and others 2015). Root biomass was sampled every fall by collecting five 2.2 cm diameter x 10 cm deep soil samples (Giddings Machine Company, Windsor, CO, USA) per plot (450 samples year-1). The samples were dried at 30 °C and roots were sorted from the sample by hand. We sorted each sample for 1 h which allowed to retrieve over 90% of all roots present in the samples (Risch and others 2013). The roots were then dried at 65 °C for 48 and weighed to the nearest mg. We averaged the values per plot and used the 2013 data only in this study.
Sampling of edaphic communities
In 2009, 2010, and 2011 we collected three composited soil samples (5 cm diameter x 10 cm depth; AMS Samplers, American Falls, ID, USA) and assessed bacterial community structure using T-RFLP profiling (Liu and others 1997; Blackwood and others 2003; Hodel and others 2014). We detected a total of 89 operational taxonomic units (OTUs). These values are in accordance with other studies reporting OTU richness (Wirthner and others 2011; Zumsteg and others 2012; Meola and others 2014) using T-RFLP profiling, a method that detects the most abundant, and thus likely, the most relevant, taxa. We averaged the data over the three years of collections for our calculations. Microbial biomass carbon (MBC) was determined with the substrate-induced method (Anderson and Domsch 1978) every fall (September) between 2009 and 2013 by collecting three mineral soil samples (5 cm diameter × 10 cm mineral soil core, AMS Samplers, American Falls, ID, USA). The three samples were combined (90 samples for each sampling year), immediately put on ice, taken to the laboratory, passed through a 2-mm sieve and stored at 4°C. Again, we only used the 2013 data in this study.
Soil samples (5 cm diameter x 10 cm depth) to extract soil arthropods were collected in June, July, and August 2011 with a soil corer lined with a plastic sleeve to ensure an undisturbed sample (total of 270 samples). The plastic line core was immediately sealed on both ends using cling film and put into a cooler. All plots were sampled within three days and the extraction of arthropods started the evening of the sampling day using a high-gradient Tullgren funnel apparatus (Crossley and Blair 1991; Vandegehuchte and others 2015). Samples were kept in the extractor for four days and the soil arthropods were collected in 95% ethanol. All individuals were counted and each individual was identified morphologically to the lowest level feasible [76 taxa, including orders, suborders, subfamilies, families (Protura, Thysanoptera, Aphidina, Psylina, Coleoptera, Brachycera, Nematocera, Auchenorryncha, Heteroptera, Formicidae); sub-phylum for Myriapoda, for Acari and Collembola also including morpho-species). Note that we also included larval stages (nine of the 76 taxa) (Vandegehuchte and others 2015). All data were summed over the season. A detailed species list for mites and collembolans is published (Vandegehuchte and others 2017a) [https://doi.org/10.1371/journal.pone.0118679.s001]. Earthworms are rare in the SNP and therefore were not included. We collected eight random 2.2 cm diameter x 10 cm deep soil cores from each plot in September 2013 to determine the soil nematode community composition. The samples were mixed and the nematodes were extracted from 100 ml of fresh soil using Oostenbrink elutriators (Oostenbrink 1960). All nematodes in a 1 ml of the 10 ml extract were counted, a minimum of 150 individuals sample-1 were identified to genus or family level using (Bongers 1988), the numbers of all nematodes were extrapolated to the entire sample and expressed for a 100 g dry sample. In total we identified 63 genus or family levels (Vandegehuchte and others 2015). The list of all the nematodes found is published (Vandegehuchte and others 2015) [http://www.oikosjournal.org/appendix/oik-03341] or DOI: [doi: 10.1111/oik.03341].
We are aware that sampling soil microbes from 2009 to 2011 and soil arthropods in 2011 was not ideal, but we are positive that this does not bias the results. Most of the parameters measured in our experiment either already showed a treatment response after the first growing season (e.g., plant biomass) or did not respond over the entire time experiment (e.g., microbial biomass C). The microbial community composition (2009 – 2011) was highly influenced by inter-annual differences in temperature and precipitation, but did not differ between treatments or vegetation types (Hodel and others 2014). We therefore felt comfortable using the 2009 through 2011 data for describing the soil microbial community in our experimental treatments. Similarly, we are positive that our soil arthropod data are representative. We did assess soil arthropods in August 2012 and found no differences to the August 2011 data. However, we did not feel comfortable combining the 2011 June, July, August data with only August data for 2012 for our analyses.
Sampling of soil properties
We collected three soil samples (5 cm diameter x 10 cm depth) in each plot in September 2013 after removing the vegetation. First, we collected the top layer of mineral soil rich in organic matter, the surface organic layer or rhizosphere, typically 1 to 3 cm in depth with a soil corer (AMS Samples, American Falls, Idaho, USA). Second, we collected a 10 cm mineral soil core beneath this surface layer. The cores for each layer were composited, dried at 65 °C for 48 h and fine-ground to pass a 0.5 mm screen. We then analysed all samples for total C using a Leco TruSpec Analyser (Leco, St. Joseph, Michigan, USA). Mineral soil pH was measured potentiometrically in 1:2 soil:CaCl2 solution with an equilibration time of 30 min.
Soil net N mineralisation was assessed during the 2013 growing season (Risch and others 2015). For this purpose, we randomly collected a 5 cm diameter x10 cm deep soil sample with a soil corer (AMS Samples, American Falls, Idaho, USA) after clipping the vegetation in June 2013. After weighing and sieving (4 mm mesh) the soil, we extracted a 20 g subsample in 1 mol l-1 KCl for 1.5 h on an end-over-end shaker and thereafter filtered it through ashless folded filter paper (DF 5895 150, ALBET LabScience, Hahnenmühle FineArt GmbH, Dassel, Germany). From these filtrates NO3- concentrations were measured colorimetrically (Norman and Stucki 1981) and NH4+with flow injection analysis (FIAS 300, Perkin Elmer, Waltham Massachusetts, USA) (Risch and others 2015). We dried the rest of the sample 105 °C to constant mass to determine fine,fraction bulk density. A second soil sample was collected within each plot in June 2013 with a corer lined with a 5 x 13 cm aluminium cylinder. The corer was driven 11.5 cm deep into the soil so that the top 1.5 cm of the cylinder remained empty. Into this space we placed a polyester bag (250 µm) filled an ion-exchanger resin to capture the incoming N. The bag was filled with a 1:1 mixture of acidic and alkaline exchanger resin (ion-exchanger I KA/ion exchanger IIIAA, Merck AG, Darmstadt, Germany). We then removed 1.5 cm soil at the bottom of the cylinder and placed a second resin exchanger bag into this space to capture the N leached from the soil column. To assure that the exchange resin was saturated with H+ and Cl- prior to filling the bags, the mixture was stirred with 1.2 ml l-1 HCl for 1 h and then rinsed with demineralized water until the electrical conductivity of the water reached 5 µm cm-1. The cylinder with the resin bags in place was reinserted into the soil with the top flush to the soil surface and incubated for three months. We recollected the cylinders in September 2013. Each resin bag and 20 g of sieved soil (4 mm mesh) from each cylinder were then separately extracted with KCl and NO3- and NH4+ concentrations were measured. Nitrate and NH4+ concentrations of all samples were then converted to a content basis by multiplying their values with fine fraction bulk density. Net N mineralisation was thereafter calculated as the difference between the N content of the samples collected at the end of the three-month incubation (including the N extracted from the bottom resin bag) and the N content at the beginning of the incubation (Risch and others 2015).
Soil CO2 emissions were measured every two weeks between 0900 and 1700 hrs from early May through late September 2013 with a PP-Systems SRC-1 soil respiration chamber (15 cm high, 10 cm diameter; closed circuit) attached to a PP-Systems EGM-4 infrared gas analyser (PP-Systems, Amesbury, MA, USA) on two locations per plot (Risch and others 2013). The chamber was placed on randomly placed, permanently installed PVC collars (10 cm diameter) driven 5 cm into the soil at the beginning of the study (Risch and others 2013). Freshly germinated plants growing within the collars were removed prior to each measurement to avoid measuring plant respiration or photosynthesis. The two measurements collected per plot and sampling date were averaged.
Soil moisture (with time domain reflectometry; Field-Scout TDR-100, Spectrum Technologies, Plainfield, Illionois, USA) and temperature (with a waterproof digital pocket thermometer; Barnstead International, Dubuque, Iowa, USA) were measured at five random locations per plot every two weeks during the growing seasons during the experiment for the 0 to 10 cm depth (Risch and others 2013, 2015). As soil moisture and soil temperature were highly negatively correlated (Risch and others 2013), we only used soil moisture for this study. We used plot-level averages of all values available to capture soil moisture variability during the five years of the experiment. The results remained unchanged when we only used soil moisture from the 2013 growing season.
Numeral calculations and statistical analyses
Ecosystem coupling. We conducted principal component analyses (PCAs; unscaled) at the complete dataset level using the abundances of each taxonomical entity to describe each of the five different communities used in this study: aboveground-dwelling invertebrates, vascular plants, soil microorganisms, soil arthropods and soil nematodes. We retained the first two components (PCA axis 1 and PCA axis 2) of each analysis as we found them to adequately represent the temporal and spatial variability of our 90 treatment plots in previous studies55,67. Together they explained a total of 71.70% of the variation for aboveground invertebrates, 44.36% for plants, 44.85% for soil microorganisms, 61.85% for soil arthropods and 77.19% for soil nematodes. In addition, we used soil pH and soil organic C content as a proxy for soil chemical properties, soil bulk density as a proxy for soil physical properties and soil moisture (negatively correlated with soil temperature) as a proxy for soil micro-climatic conditions for an overall total of fourteen constituents.
We calculated ecosystem coupling9 for each exclosure treatment within each vegetation type (i.e., 2  5 treatment combinations in total) as an integrated measure of pairwise ecological interactions between ecosystem constituents representing ecological communities and the soil abiotic environment. These ecological interactions are defined by non-parametric Spearman rank correlation analyses between two constituents, excluding interactions involving two abiotic constituents (e.g., soil pH vs. soil moisture) and interactions between the first (PC1) and second (PC2) component of each community type, as these are orthogonal by definition. Interactions between abiotic constituents were excluded from the analyses because the focus of our study was on communities and how they interact with one another and their surrounding environment; therefore, including abiotic-abiotic interactions was not of interest here. Given that the effectiveness of our experimental design resulted in that no community composition data of aboveground-dwelling invertebrates was available for the “None” plots (all animals excluded), only thirteen instead of fourteen constituents were included in the ecosystem coupling calculations for this treatment. The complete absence of aboveground invertebrates represents the most extreme case of disturbance between aboveground animal communities and the rest of the ecosystem constituents. This may have resulted in a slight overestimation of ecosystem coupling for these plots.
	Average ecosystem coupling was calculated as follows:
Ecosystem coupling=
where Xi is the absolute Coupling was calculated value of the Spearman’s rho coefficient of the ith correlation for each treatment within each vegetation type (i.e., based on nine replicates each), considering and n is the number of pairwise comparisons considered (n = a total of 80; interactions (56 in the case of the “None” treatment). We considered a total of 40 biotic-biotic interactions (i.e., concerning two community-level principal components such as plants and microbes; 24 in the case of the “None” treatment) and 40 abiotic-biotic (i.e., concerning one community-level principal component and one abiotic factor, e.g., plant community and soil properties; 32 in the case of the “None” treatment).	Coupling was calculated for each treatment within each vegetation type (i.e., based on nine replicates each), considering a total of 80 interactions (56 in the case of the “None” treatment). We considered a total of 40 biotic-biotic interactions (i.e., concerning two community-level principal components such as plants and microbes; 24 in the case of the “None” treatment) and 40 abiotic-biotic (i.e., concerning one community-level principal component and one abiotic factor, e.g., plant community and soil properties; 32 in the case of the “None” treatment).
To establish whether constituents were significantly and positively coupled within treatments (i.e., the average of their correlation coefficients were greater than in a null model where correlation only happens by chance), we calculated one-tailed p-values based on permutation tests with 999 permutations.

We considered six ecosystem functions and process rates commonly used to assess ecosystem functioning (Meyer and others 2015; Manning and others 2018). Plant N content represents a measure of forage quality, while plant richness has been shown to stabilise biomass production, thus allowing the system to respond to changes in herbivory. Soil net N mineralisation, soil respiration, root biomass, and microbial biomass represent fluxes or stocks of energy. For all functions and processes higher values represent higher functioning (Manning and others 2018). All these variables were measured in the last year of the experiment (2013). We then quantified ecosystem multifunctionality using the multiple threshold approach (Byrnes and others 2014; Manning and others 2018), which considers the number of functions that are above a certain threshold, over a series of threshold values (typically 10-99%) that are defined based on the maximum value of each function. We weighted all our functions equally for these calculations (Manning and others 2018). The number of functions in a plot with values higher than a given threshold value for the respective function is summed up. The sum represents ecosystem multifunctionality for that plot. Given that choosing any particular threshold as a measure of ecosystem multifunctionality is arbitrary, we calculated the average of thresholds from 10-90% (in 10% intervals) as a more integrated representation of ecosystem multifunctionality.
We used Pearson correlations to explore the relationships between ecosystem coupling (all interactions, biotic-biotic interactions, abiotic-biotic interactions involving above- and belowground constituents, and all interactions, biotic-biotic interactions, abiotic-biotic interactions involving belowground constituents only) and ecosystem multifunctionality by calculating the slopes of all relationships between ecosystem coupling and multifunctionality for all thresholds between 10 and 99%. We also related ecosystem coupling with the average of multifunctionality at thresholds between 30-80% as explained before and considered this correlation as a robust indication of the type of association between these two variables. In addition, we explored the relationships between ecosystem coupling (all interactions, biotic-biotic interactions, abiotic-biotic interactions involving above- and belowground constituents, and all interactions, biotic-biotic interactions, abiotic-biotic interactions involving belowground constituents only) and individual ecosystem functions. The effects of exclosures and vegetation type on individual functions and multifunctionality were evaluated using linear mixed effects models ('lme' function of the nlme package), with exclosure and vegetation type as fixed effects and fence as a random factor. All statistical analyses and numerical calculations were done in R version 3.4.0 (R Core Team 2016).

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- R Core Team. 2016. R: A language and environment for statistical computing. Vienna, Austria: R Foundation for Statistical Computing
- Risch AC, Haynes AG, Busse MD, Filli F, Schütz M. 2013. The response of soil CO2 fluxes to progressively excluding vertebrate and invertebrate herbivores depends on ecosystem type. Ecosystems 16:1192–202.
- Risch AC, Schütz M, Vandegehuchte ML, Van Der Putten WH, Duyts H, Raschein U, Gwiazdowicz DJ, Busse MD, Page-Dumroese DS, Zimmermann S. 2015. Aboveground vertebrate and invertebrate herbivore impact on net N mineralization in subalpine grasslands. Ecology 96:3312–22.
- Schütz M, Risch AC, Achermann G, Thiel-Egenter C, Page-Dumroese DS, Jurgensen MF, Edwards PJ. 2006. Phosphorus translocation by red deer on a subalpine grassland in the Central European Alps. Ecosystems 9:624–633.
- Schütz M, Risch AC, Leuzinger E, Krüsi BO, Achermann G. 2003. Impact of herbivory by red deer (Cervus elaphus L.) on patterns and processes in subalpine grasslands in the Swiss National Park. For Ecol Manage 181:177–88.
- Vandegehuchte ML, van der Putten WH, Duyts H, Schütz M, Risch AC. 2017a. Aboveground mammal and invertebrate exclusions cause consistent changes in soil food webs of two subalpine grassland types, but mechanisms are system-speciﬁc. Oikos 126:212–23.
- Vandegehuchte ML, Raschein U, Schütz M, Gwiazdowicz DJ, Risch AC. 2015. Indirect short- and long-term effects of aboveground invertebrate and vertebrate herbivores on soil microarthropod communities. PLoS One 10:e0118679.
- Vandegehuchte ML, Schütz M, de Schaetzen F, Risch AC. 2017b. Mammal-induced trophic cascades in invertebrate food webs are modulated by grazing intensity in subalpine grassland. J Anim Ecol 86:1434–46.
- Vandegehuchte ML, Trivellone V, Schütz M, Firn J, de Schaetzen F, Risch AC. 2018. Mammalian herbivores affect leafhoppers associated with specific plant functional types at different timescales. Funct Ecol 32:545–55.
- Wirthner S, Frey B, Busse MD, Schütz M, Risch AC. 2011. Effects of wild boar (Sus scrofa L.) rooting on the bacterial community structure in mixed-hardwood forest soils in Switzerland. Eur J Soil Biol 47:296–302. http://dx.doi.org/10.1016/j.ejsobi.2011.07.003
- Zumsteg A, Luster J, Göransson H, Smittenberg RH, Brunner I, Bernasconi SM, Zeyer J, Frey B. 2012. Bacterial, Archaeal and Fungal Succession in the Forefield of a Receding Glacier. Microb Ecol 63:552–64. https://doi.org/10.1007/s00248-011-9991-8

ecosystem-coupling

display_name: ecosystem-coupling

ecosystem-functions

display_name: ecosystem-functions

ecosystem-multifunctionality

display_name: ecosystem-multifunctionality

exclusion

display_name: exclusion

grassland

display_name: grassland

grazing

display_name: grazing

invertebrates

display_name: invertebrates

subalpine

display_name: subalpine

swiss-national-park

display_name: swiss-national-park

vertebrates

display_name: vertebrates

education

en: Education and science

envidat

en: EnviDat: the Environmental Data Portal

en: EnviDat is the WSL data portal providing unified and managed access to environmental monitoring and research data. The portal has the capability to host and publish data sets. Through the sharing of unique and high-value data sets worldwide, EnviDat fosters research and contributes to advancing environmental science.

en: Ecosystem coupling and multifunctionality - exclosure experiment


maintainer: EnviDat Support

en: wind-velocity wind-direction wind-speed anemometer wind raclets

state: active

type: dataset

en: Sonic time

media_type: npy

en: The data set contains the time in YYYY-mm-dd HH:MM:SS.sss for the anemometer data. The time is given in UTC+1.

format: SERVICE

en: Sonic wind component w

en:
The data contains values for w direction of the wind in a sampling frequency of 20 Hz.
Erroneour spikes were filterd in two steps.

1. Define moving window uₓ = u(i-3:i+3)
2. If u(i)-mean(uₓ) > 2*std(uₓ):u(i) = NaN

Frozen detectors resulted in continuous zero values (not filtered).
The wind speed and direction were calculated as follows:

wind speed = sqrt(u² + v²)
wind direction = 180° + (180° / PI)* arctan2 (-u, -v)

en: Sonic temperature

en: The data set contains values of temperature (°C) measured with the anemometer at a sampling frequency of 20 Hz.

en: Sonic wind component v

en:
The data contains values for v direction of the wind in a sampling frequency of 20 Hz.
Erroneour spikes were filterd in two steps.

1. Define moving window uₓ = u(i-3:i+3)
2. If u(i)-mean(uₓ) > 2*std(uₓ):u(i) = NaN

Frozen detectors resulted in continuous zero values (not filtered).
The wind speed and direction were calculated as follows:

wind speed = sqrt(u² + v²)
wind direction = 180° + (180° / PI)* arctan2 (-u, -v)

en: Sonic wind component u

en:
The data contains values for u direction of the wind in a sampling frequency of 20 Hz.
Erroneour spikes were filterd in two steps.

1. Define moving window uₓ = u(i-3:i+3)
2. If u(i)-mean(uₓ) > 2*std(uₓ):u(i) = NaN

Frozen detectors resulted in continuous zero values (not filtered).
The wind speed and direction were calculated as follows:

wind speed = sqrt(u² + v²)
wind direction = 180° + (180° / PI)* arctan2 (-u, -v)

en: A Young 81000 sonic anemomenter was deployed at Gotschnagrat (LON: 46.859 LAT: 9.849) to record three components of the wind velocity (u, v, w in [m s‾ ¹]) and air temperature (Ts in [°C]). The anemomenter was mounted in direction North at a height of 1.5 m above snow surface at the beginning. The time within each data set is given in UTC+1. Instrument specifications can be found [here](http://www.youngusa.com/Manuals/81000-90(I).pdf) .

anemometer

display_name: anemometer

raclets

display_name: raclets

wind

display_name: wind

wind-direction

display_name: wind-direction

wind-speed

display_name: wind-speed

wind-velocity

display_name: wind-velocity

education

fr: Education, science

de: Bildung, Wissenschaft

it: Formazione e scienza

fr: EnviDat: le portail de données environnementales

de: EnviDat: das Umwelt-Datenportal

it: EnviDat: il portale dei dati ambientali

fr: EnviDat est le portail de données du WSL; il offre un accès uniformisé et réglementé aux données de surveillance et de recherche en matière d’environnement. Ce portail est capable d’héberger et de publier des ensembles de données précieuses et uniques, qui sont partagées aux quatre coins de la planète. Ce faisant, EnviDat favorise la recherche et contribue à l’avancement des sciences environnementales.

de: EnviDat ist das Datenportal der WSL für einen einheitlichen und verwalteten Zugang zu Umweltdaten aus Monitoring- und Forschungsprojekten. Auf dem Portal können Datensätze gehostet und veröffentlicht werden. Durch die weltweite Bereitstellung einzigartiger und wertvoller Datensätze leistet EnviDat einen Beitrag zur Forschung im Bereich der Umweltwissenschaften.

it: EnviDat è il portale dati del WSL che fornisce un accesso unificato e gestito ai dati di monitoraggio ambientale e di ricerca. Il portale ha la capacità di ospitare e pubblicare serie di dati preziosi e unici che vengono condivisi in tutto il mondo. Così facendo, EnviDat promuove la ricerca e contribuisce al progresso della scienza ambientale.

political_level: other

wsl

capacity: public

type: organization

approval_status: approved

key: guid

key: uri

value: Envidat Harvester

en: Snow Drift Station - 3D Ultrasonic


disdrometer-data-davos-wolfgang

title_for_slug: disdrometer-data-davos-wolfgang

en: precipitation fall-velocity disdrometer davos raclets

en: Disdrometer Data Davos Wolfgang

media_type: application/zip

en:
The zip folder contains one netCDF file per day for Davos Wolfgang from February 8 to March 19 2019 with a temporal resolution of 30 seconds (time in UTC) and includes:

- characteristics of hydrometeors (e.g. diameter, fall velocity, amount per diameter class,...)
- precipitation rate
- radar reflectivity
- visibility range
- weather codes
- instrument performance

format: ZIP

mimetype: application/zip

en: The dataset contains information on precipitation amount and type for Davos Wolfgang (LON: 9.853594, LAT: 46.835577) from February 8 to March 19 2019. It includes: characteristics of hydrometeors (e.g. diameter, fall velocity, amount per diameter class,...), precipitation rate, radar reflectivity, visibility range, weather codes and instrument performance.

davos

display_name: davos

disdrometer

display_name: disdrometer

fall-velocity

display_name: fall-velocity

precipitation

display_name: precipitation

education

en: Education and science


maintainer: Jana von Freyberg

en: review uncertainties stable-water-isotopes ecohydrology plant-water-uptake root-zone endmember-mixing-models

en: Data set of MEE-20-04-264

media_type: application/pdf

en: The following two tables contain information about the data sources of the values reported in Table 1 and 2 in the paper “Plant and root-zone water isotopes are difficult to measure, explain, and predict: some practical recommendations for determining plant water sources” published in the journal ‘Methods in Ecology and Evolution’.

format: PDF

mimetype: application/pdf

en: Data set of MEE-20-04-264 (CSV)

media_type: CSV

format: CSV

en: The following two tables contain information about the data sources of the values reported in Table 1 and 2 in the paper “Plant and root-zone water isotopes are difficult to measure, explain, and predict: some practical recommendations for determining plant water sources” published in the journal 'Methods in Ecology and Evolution'.

ecohydrology

display_name: ecohydrology

endmember-mixing-models

display_name: endmember-mixing-models

plant-water-uptake

display_name: plant-water-uptake

review

display_name: review

root-zone

display_name: root-zone

stable-water-isotopes

display_name: stable-water-isotopes

uncertainties

display_name: uncertainties

envidat

en: EnviDat: the Environmental Data Portal

en: EnviDat is the WSL data portal providing unified and managed access to environmental monitoring and research data. The portal has the capability to host and publish data sets. Through the sharing of unique and high-value data sets worldwide, EnviDat fosters research and contributes to advancing environmental science.

en: Data set of: Plant and root-zone water isotopes are difficult to measure, explain, and predict: some practical recommendations for determining plant water sources


maintainer: Anita C. Risch

en: soil-chemistry nutrient-network global grassland worldwide soil soil-biology mineralization

en:
site.code: Site code

Continent: Continent

sampleID: Sample ID

nn.latitude: Lattitude

nn.longitude: Longitude

bioclim.T.ann: Mean annual temperature (°C)

bioclim.T.var: Temperature variability (°C)

bioclim.T.q.wet: Temperature of the wettest quarter (°C)

bioclim.T.q.dry: Temperature of the driest quarter (°C)

bioclim.T.q.warm: Temperature of the warmest quarter (°C)

bioclim.P.ann: Mean annual preciptiation (mm)

bioclim.P.q.wet: Precipitation of the wettest quarter (mm)

ch.pH: pH

ch.WSK.vol: Water holding capacity (vol%)

ch.bulkdensity.gcm3: Bulk density (g cm-3)

ch.porespace: Pore space (vol%)

ch.sand.perc: Soil texture: sand (%)

ch.silt.perc: Soil texture: silt (%)

ch.clay.perc: Soil texture: clay (%)

ch.NO3start.mgg: NO3 at the beginning of the incubation (mgkg)

ch.Ntot.perc: Total soil N (%)

ch.Ctot.perc: Total soil C (%)

ch.Corg.perc: Soil organic C (%)

ch.netNminfield.mgkgd: Realised soil net N mineralisation (mg kg-1 d-1)

ch.netNminlab.mgkgd: Potential soil ne N mineralisation (mg kg-1 d-1)

archaea.cpg.log: Ammonia oxidising archaea (archaeal amoA gene copies g-1 soil)

bacteria.cpg.log: Ammonia oxidising bacteria (bacterial amoA gene copies g-1 soil)

micr.biom: Microbial biomass (ug g-1 soil)

ch.CtoN: C to N ratio

ch.netNnitriffield.mgkgd.sqrt: Realised soil net nitrification, transformed (mg kg-1 d-1)

ch.netNammoniffield.mgkgd.sqrt: Realised soil net ammonification, transformed (mg kg-1 d-1)

ch.netNnitriflab.mgkgd.sqrt: Potential soil net nitrification, transformed (mg kg-1 d-1)

ch.netNammoniflab.mgkgd.sqrt: Potential soil net ammonifiation, transformed (mg kg-1 d-1)

format: XLS

en:
This dataset contains all data on which the following publication below is based.

Paper Citation:

Risch, A. C.; Zimmermann, S.; Ochoa-Hueso, R.; Schütz, M.; Frey, B.; Firn, J. L.; Fay, P. A.; Hagedorn, F.; Borer, E. T.; Seabloom, E. W.; et al. Soil net nitrogen mineralisation across global grasslands. Nat. Commun. 2019, 10 (1), 4981 (10 pp.). doi.org/10.1038/s41467-019-12948-2

Please cite this paper together with the citation for the datafile.

We conducted coordinated measurements of realised and potential soil net Nmin, and assessed water holding capacity, bulk density, C and N content, texture, pH, pore space, microbial biomass, and archaeal (AOA) and bacterial (AOB) ammonia oxidiser abundance using identical materials and methods across 30 grasslands on six continents. The sites covered a globally relevant range of climatic and edaphic conditions. Climate data was obtained from worldclim - Global climate data https://www.worldclim.org/

global

display_name: global

grassland

display_name: grassland

mineralization

display_name: mineralization

nutrient-network

display_name: nutrient-network

soil

display_name: soil

soil-biology

display_name: soil-biology

soil-chemistry

display_name: soil-chemistry

worldwide

display_name: worldwide


en: x-ray-tomography microstructure snow-mechanics compression-experiments snow

en: Snowmicroquakes Data

en: Compression experiments, X-ray images and simulation data with supplemented README file

en: README

media_type: TXT

en: README file explaining the organization of the data

format: TXT

en:
When snow is compressed with a certain speed, micro-snowquakes are triggered in the porous structure of bonded crystals. The present dataset covers uniaxial compression experiments of snow at different strain rates and concurrent X-ray tomography imaging documenting this feature.

The experiments were conducted in a micro-compression stage operated in the X-ray tomography scanner in the SLF cold laboratory. The dataset comprises the compression force data of 17 compression experiments, the 3D image data from 4 X-ray tomography scans and the results of numerical simulations.

compression-experiments

display_name: compression-experiments

microstructure

display_name: microstructure

snow

display_name: snow

snow-mechanics

display_name: snow-mechanics

x-ray-tomography

display_name: x-ray-tomography

en: Compressive stick slip and snow-micro-quakes


snow-drift-station-micro-rain-radar

title_for_slug: snow-drift-station-micro-rain-radar

en: raclets snow radar mountain precipitation rain

en: MRR data of period 3

en: The dataset contains measurements between March 8 and March 13, 2019, with time in UTC. The parameters and specifications are given in MRR_parameters.pdf. The zip folder contains hourly data files in netCDF format and a log file, reporting the instrument status.

en: Parameter MRR

en: The PDF contains additional information on parameters which were set for the certain time periods (time in UTC).

en: MRR data of period 4

en: The dataset contains measurements between March 13 and March 22, 2019, with time in UTC. The parameters and specifications are given in MRR_parameters.pdf. The zip folder contains hourly data files in netCDF format and a log file, reporting the instrument status.

en: MMR Data Period 1

en: The dataset contains measurements between March 1 and March 5, 2019, with time in UTC. The parameters and specifications are given in MRR_parameters.pdf. The zip folder contains hourly data files in netCDF format and a log file, reporting the instrument status.

en: MRR data Period 2

en: The dataset contains measurements between March 5 and March 8, 2019, with time in UTC. The parameters and specifications are given in MRR_parameters.pdf. The zip folder contains hourly data files in netCDF format and a log file, reporting the instrument status.

en: The instrument (MRR, Metek) was mounted at Gotschnagrat (LON: 46.859 LAT: 9.849) at a height of 1 m above snow surface (at the beginning of the campaign) with an orientation of 22° with respect to North and a horizontal viewing direction. The sampling time was either 5 s or 10 s, depending on the settings at the specific period. The MRR produces standard outputs like radar reflectivity, doppler velocity, etc., and additional information can be found [here](https://metek.de/de/product/mrr-2/).

mountain

display_name: mountain

precipitation

display_name: precipitation

raclets

display_name: raclets

radar

display_name: radar

rain

display_name: rain

education

en: Education and science

en: Snow Drift Station - Micro Rain Radar


en: outgoing shortwave-radiation longwave-radiation incoming davos

en: Parameter file

media_type: text/plain

mimetype: text/plain

en: Metadata

en: Processing doc

en: DATA ACCESS

media_type: SMET

en:
Incoming and outgoing shortwave and longwave 2 min radiation measurements in Davos Dorf, CH.

 References
1. Marty, C., Philipona, R., Frohlich, C., Ohmura, A.. Altitude dependence of surface radiation fluxes and cloud forcing in the alps: results from the alpine surface radiation budget network. 2002. Theoretical and Applied Climatology. Volume 72. Issue 3-4. 137-155. http://dx.doi.org/10.1007/s007040200019. 10.1007/s007040200019.
2. Christoph Marty. Surface Radiation, Cloud Forcing and Greenhouse Effect in the Alps. 2000. Institute fuer Klimaforschung ETH. Zuercher Klima-Schriften. Volume 79. http://e-collection.library.ethz.ch/eserv/eth:23491/eth-23491-01.pdf.

davos

display_name: davos

incoming

display_name: incoming

longwave-radiation

display_name: longwave-radiation

outgoing

display_name: outgoing

shortwave-radiation

display_name: shortwave-radiation

envidat

en: EnviDat: the Environmental Data Portal

en: EnviDat is the WSL data portal providing unified and managed access to environmental monitoring and research data. The portal has the capability to host and publish data sets. Through the sharing of unique and high-value data sets worldwide, EnviDat fosters research and contributes to advancing environmental science.

en: ASRB_DAV: Shortwave and longwave radiation measurements (2 min) in Davos Dorf


maintainer: Roksolana Petrovska

en: regeneration acer mortality-rate dominance fagus lar shade-tolerance

media_type: r script (flat-ascii text)

en: The R-script of survival analysis for accelerated failure time model

en: Mortality of regeneration

en: Acer platanoides, Acer pseudoplatanus and Fagus sylvatica living and dead juveniles (0-500 cm in height) grown in the deep shade of a prime beech-dominated forest.

en:
 One individual per species, vitality class (low and high) and height class (eight classes: 0–10, 11–20, 21–35, 36–60, 61–90, 91–130, 131–200 and 201–500 cm) was randomly selected and harvested in each of the six plots. This resulted in a sample of 82, 80 and 89 living individuals of A. platanoides, A. pseudoplatanus and F. sylvatica, respectively.

 Additionally stems of dead Acer spp. and F. sylvatica trees that had died within the last three years (2015–2018) were randomly harvested, matching the height classes of the harvested living trees wherever possible. In total, 179 dead young trees (60 A. platanoides, 72 A. pseudoplatanus and 47 F. sylvatica) were collected.

 Variables:
* species_code: a_pla - Acer platanoides, a_pse - Acer pseudoplatanus, f_syl - Fagus sylvatica
* species: as above
* dummy: 0 - living individual, 1 - dead individual
* LAR_cm2_g: leaf area ratio or ratio of leaf area to total plant biomass, [cm2/g]
* tree_age: in years
* avg_ring_micron: average width of the last 5 rings in tree life excluding the last ring
* dry_mass_g: aboveground and belowground biomass
* DLI: direct light index (measured only under living individuals)
* BLI: diffuse light index (measured only under living individuals)
* GLI: global light index

acer

display_name: acer

dominance

display_name: dominance

fagus

display_name: fagus

lar

display_name: lar

mortality-rate

display_name: mortality-rate

regeneration

display_name: regeneration

shade-tolerance

display_name: shade-tolerance

en: Mortality of regeneration: Acer spp. and Fagus sylvatica


en: snow-distribution snow-depth canopy-height forest-structure airborne-laser-scanning

en: Canopy height maps

en: Canopy height maps of each of the three field areas Davos, Engadine and Grand Mesa

en: Readme

en: Snow depth maps

en: Snow depth maps from two snow-on flights at each of the three field areas Davos (March 20th and 31st 2017), Engadien (March 20th and 31st, 2017) and Grand Mesa (February 8th and 16th, 2017)

en: Terrain elevation maps

en: Digital elevation model computed from snow-off flights for each of the field areas Davos, Engadine and Grand Mesa

en:
This dataset includes snow depth, canopy height and terrain elevation maps of forest stands in the Grisons (CH) and at Grand Mesa (CO,USA) derived from airborne lidar. Data were acquired i) within a pilot mission of NASA's Airborne Snow Observatory in the Swiss Alps in March 2017 and ii) during NASA’s SnowEx campaign at Grand Mesa in February 2017. Snow depth maps are available for two dates separated by approx.1 week, and include an area of ca. 0.5km2 for each of the three sites Davos, Engadine and Grand Mesa.
All data were presented and analyzed in the publication 'Revisiting Snow Cover Variability and Canopy Structure within Forest Stands: Insights from Airborne Lidar Data' (Mazzotti et al., 2019, WRR, doi: 10.1029/2019WR024898). This publication must be cited when using this dataset.

Paper Citation:
 > _Giulia Mazzotti; William Ryan Currier; Jeffrey S. Deems; Justin M. Pflug; Jessica D. Lundquist; Tobias Jonas (2019). Revisiting Snow Cover Variability and Canopy Structure Within Forest Stands: Insights From Airborne Lidar Data. Water Resources Research, 55, 6198– 6216, [doi: 10.1029/2019WR024898](https://doi.org/10.1029/2019WR024898)._

airborne-laser-scanning

display_name: airborne-laser-scanning

canopy-height

display_name: canopy-height

forest-structure

display_name: forest-structure

snow-depth

display_name: snow-depth

snow-distribution

display_name: snow-distribution

education

en: Education and science

en: ALS-based snow depth and canopy height maps from flights in 2017 (Grisons, CH and Grand Mesa, CO)


en: long-term-experiment mountain-forest treeline afforestation stillberg

en: Site photo

media_type: JPEG

en: This site photo is outdated and has been replaced with an updated version (see description).

en: Documentation

media_type: DOC

en: This data documentation is outdated and has been replaced with an updated version (see description).

mimetype: application/msword

en: Dataset V1

media_type: GDB

en: This dataset is outdated and has been replaced with an updated version (see description).

format: GDB

en:
 Important
This EnviDat entry is outdated. The most recent, usable version of the data can be found under the new EnviDat entry "Long-term afforestation experiment at the Alpine treeline site Stillberg, Switzerland." The entry can be found under this link <https://www.envidat.ch/dataset/afforestation-stillberg> and with this DOI 10.16904/envidat.397.

afforestation

display_name: afforestation

long-term-experiment

display_name: long-term-experiment

mountain-forest

display_name: mountain-forest

stillberg

display_name: stillberg

treeline

display_name: treeline

envidat

en: EnviDat: the Environmental Data Portal

en: EnviDat is the WSL data portal providing unified and managed access to environmental monitoring and research data. The portal has the capability to host and publish data sets. Through the sharing of unique and high-value data sets worldwide, EnviDat fosters research and contributes to advancing environmental science.

en: Long-term treeline research dataset at Stillberg, Davos


maintainer: Stephan Zimmermann

en: magnesium biogeochemical-cycling potassium calcium fagus-orientalis-lipsky

en: Concentrations of Ca, Mg, K, and Na in throughfall, litterflow and soil solution of a Hyrcanian forst in Northern Iran

en:
Throughfall, litterflow and soil solution were sampled during one whole year under five Oriental beech trees in a mixed Hyrcanian beech forest. The amounts of Ca2+, Mg2+, K+ and Na+ in these fluxes were calculated based on their concentrations and the sampled volumes, and subsequently compared with the respective fluxes in the rainfall and soil solution of an adjacent forest gap.
In addition six soil profiles, one close to every single tree and one in the forest gap, were analyzed for pH, CaCO3, organic matter and texture.

biogeochemical-cycling

display_name: biogeochemical-cycling

calcium

display_name: calcium

fagus-orientalis-lipsky

display_name: fagus-orientalis-lipsky

magnesium

display_name: magnesium

potassium

display_name: potassium

en: Base cation dynamics in an Oriental beech forest


backward-trajectories

maintainer: Michael Sprenger

title_for_slug: backward-trajectories

en: alps raclets backward-trajectories lagrangian-analysis complex-topopgraphy atmospheric-dynamics cosmo

en: Backward Trajectories

en: The trajectories are started from surface up to 5000 m altitude, with an equidistant vertical spacing of 200 m, and calculated 120 h backward in time. As long as the trajectories stay in the COSMO1 domain (essentially the Alpine region), as run operationally at MeteoSwiss, the COSMO analysis wind fields are used. When the trajectories leave this domain, they are extended further backward in time based on ECMWF analysis data. The following fields are traced along the trajectories: longitude, latitude, geopotential height, pressure, specific humidity, temperature and surface topography. Furthermore, it is indicated for each trajectory time instance on which model data, COSMO or ECMWF, the trajectory time is based. Finally, the trajectories are started every hour in the specified time period, and positions and fields are available every 10 min. Each day is saved as a separate file. The last 8 digits of the file names indicate the starting date of the trajectories.

en: Backward trajectories were calculated from two positions: Davos Wolfgang (LON: 9.85361, LAT: 46.83551) and Weissfluhjoch (LON: 9.80646 LAT: 46.83304) for the time period February 2 until March 27 2019 using COSMO or ECMWF, respectively.

alps

display_name: alps

atmospheric-dynamics

display_name: atmospheric-dynamics

backward-trajectories

display_name: backward-trajectories

complex-topopgraphy

display_name: complex-topopgraphy

cosmo

display_name: cosmo

lagrangian-analysis

display_name: lagrangian-analysis

raclets

display_name: raclets

education

en: Education and science

en: Backward  Trajectories


maintainer: Sven-Eric Hopf

en: iap fraxinus forest ash-dieback monitoring

en: Monitoring of ash trees as part of the Intercantonal Forest Observation Programme

media_type: .pdf

en: Packgroun information on data for the publication: Spread and Severity of Ash Dieback in Switzerland – Tree Characteristics and Landscape Features Explain Varying Mortality Probability, (Klesse et al. 2021, Front. For. Glob. Change 4:645920., doi: 10.3389/ffgc.2021.645920)

en:
In 2013, the Institute for Applied Plant Biology (IAP) started a monitoring programme to study the development and the spatial variation of the ash dieback disease with the aim to find some partially resistant European ash trees (Fraxinus excelsior).
We collaborate as co-authors for the publication: Spread and Severity of Ash Dieback in Switzerland - Tree Characteristics and Landscape Features Explain Varying Mortality Probability (Klesse et al. 2021 in frontiers)

ash-dieback

display_name: ash-dieback

forest

display_name: forest

fraxinus

display_name: fraxinus

iap

display_name: iap

monitoring

display_name: monitoring


maintainer: Henning Löwe

en: microstructure spherical-model-snow snow-mechanics 3d-ct-images unconfined-compression-experiments discrete-element-modelling

en: Data Structure (Python Notebook - view online)

media_type: ipynb

en: Data Access

media_type: HTTPS

en: Data Structure (Python Notebook)

en:
For the investigation of microstructural and mechanical properties of snow unconfined compression experiments and 3D computed tomography (CT) imaging were performed on sintered rounded grain snow and spherical model snow. The spherical model snow was generated to create geometrically simplified, well-defined microstructures for calibration of numerical models, such as discrete element models (DEM) in which the microstructure is represented by spherical particles.
In the experiments, microstructural variation was created by varying the sintering time (contact size) and the density of the ice sphere samples (number of contacts). The 3D CT images allow for a complete reconstruction of the entire experimental sample (cylindrical sample dimension: diameter = 33.6 mm; height = 14 mm).

discrete-element-modelling

display_name: discrete-element-modelling

microstructure

display_name: microstructure

snow-mechanics

display_name: snow-mechanics

spherical-model-snow

display_name: spherical-model-snow

unconfined-compression-experiments

display_name: unconfined-compression-experiments

envidat

en: EnviDat: the Environmental Data Portal

en: Unconfined compression experiments and 3D CT images of spherical model snow and RG snow samples


maintainer: Frank Techel

en: rutschblock-test extended-column-test avalanche-observations norway switzerland avalanche-danger

en: ECT data (Norway, Switzerland)

media_type: text/csv

mimetype: text/csv

en: Rutschblock data (Switzerland)

en: avalanche occurrence data (Norway, Sitzerland) - avalanche sizes per day and warning region

en: avalanche occurrence data (Norway, Sitzerland) - largest avalanche size per day and warning region

en:
Observational data used to quantitatively describe the key elements describing avalanche danger: snowpack stability, the frequency distribution of snowpack stability, and avalanche size.
The data set consists of
- Rutschblock test results (Switzerland)
- Extended Column Test results (Switzerland, Norway)
- Avalanche occurrence data (Switzerland, Norway).
The data were extracted from the respective operational databases of the national avalanche warning services in Switzerland (WSL Institute for Snow and Avalanche Research SLF Davos, Switzerland) and Norway (The Norwegian Water Resources and Energy Directorate NVE).
For further information regarding the data, please refer to the publication or contact the author.

avalanche-danger

display_name: avalanche-danger

avalanche-observations

display_name: avalanche-observations

extended-column-test

display_name: extended-column-test

norway

display_name: norway

rutschblock-test

display_name: rutschblock-test

switzerland

display_name: switzerland

en: EnviDat is the WSL data portal providing unified and managed access to environmental monitoring and research data. The portal has the capability to host and publish data sets. Through the sharing of unique and high-value data sets worldwide, EnviDat fosters research and contributes to advancing environmental science.

en: Stability tests, avalanche observations, Switzerland, Norway


mosaic-snow-on-sea-ice-data

title_for_slug: mosaic-snow-on-sea-ice-data

en: snow-distribution spatial-variability arctic-sea-ice surface-mass-balance drifting-and-blowing-snow

en: SNOWPACK setup for MOSAiC

media_type: various

en:
 The zip file contains both input and output data as well as the model configuration as used in the submitted manuscript "Towards a fully physical representation of snow on Arctic sea ice using a 3D snow-atmosphere model".

The zip file contains 2 main folders:
* SNOWPACK_JGR_ALPHA1
* SNOWPACK_JGR_ALPHA3

In the SNOWPACK_JGR_ALPHA1 (reference "SP_R" simulation) and SNOWPACK_JGR_ALPHA3 (comparison "SP_C" scenario) folders you find the corresponding inputs, outputs and configuration as used in the paper, as well as the settings used - which only differ by the changed "SCHMIDT_DRIFT_FUDGE" value that is found in each SNOWPACK_JGR_ALPHA/setup/io.ini file. The input data is already linked accordingly in each io.ini file.

To reproduce the results, download and compile the source code for the adjusted SNOWPACK model first, which can be obtained from https://gitlabext.wsl.ch/snow-models/snowpack.git under the "snowpack_mosaic" branch. After installing, you can run the provided model setup uploaded here.

en: ALPINE3D setup for MOSAiC

en:
The zip file contains a folder structure that must be kept as it is in order to run the simulation in the current configuration.
The setup contains both input and output data as well as the model configuration as used in the submitted manuscript
"Towards a fully physical representation of snow on Arctic sea ice using a 3D snow-atmosphere model".

The zip file contains 3 main folders:
* base_setup_files
* a3d_jgr_alpha1
* a3d_jgr_alpha3

The "base_setup_files" contains all input files that are necessary to run the reference (R) simulation ("a3d_jgr_alpha1" folder) and the comparison "C" scenario ("a3d_jgr_alpha3") folder. In the a3d_jgr_alpha1 and a3d_jgr_alpha3 folders you find the corresponding outputs as used in the paper, as well as the settings used - which only differ by the changed "SCHMIDT_DRIFT_FUDGE" value that is found in each a3d_jgr_alphax/setup/io.ini file. The input data is already linked accordingly in each io.ini file.

a3d_jgr_alpha1 also contains the detailed snow profiles for each point along the transects.

To reproduce the results, download and compile the source code for the adjusted ALPINE3D model first, which can be obtained from https://gitlabext.wsl.ch/snow-models/alpine3d.git under the "alpine3d_mosaic" branch. After installing, you can run the provided model setup uploaded here.

en:
Data accompanying David Wagners' Dissertation.
Covers model results and various input from ALPINE3D and SNOWPACK adjusted for sea ice during MOSAiC.

arctic-sea-ice

display_name: arctic-sea-ice

drifting-and-blowing-snow

display_name: drifting-and-blowing-snow

snow-distribution

display_name: snow-distribution

spatial-variability

display_name: spatial-variability

surface-mass-balance

display_name: surface-mass-balance

education

en: Education and science

en: MOSAIC Snow on Sea Ice Data


en: longwave-radiation gemstock meteo-station shortwave-radiation snow-height

en: DATA ACCESS

en:
Meteo data at Gemsstock [Level 2 dataset](http://models.slf.ch/p/dataset-processing/).

fields      = timestamp TSS HS VW DW VW_MAX OSWR ISWR ILWR RECORD AUX_VOLTAGE LOGGER_VOLTAGE LW_SENSOR_TEMP VENTILATION_FLAG OLWR

en: Meteorological station at Gemstock (3021 m asl) in Canton Uri. The station includes in/out LW/SW and a snow height sensor.

gemstock

display_name: gemstock

longwave-radiation

display_name: longwave-radiation

meteo-station

display_name: meteo-station

shortwave-radiation

display_name: shortwave-radiation

snow-height

display_name: snow-height

envidat

en: EnviDat: the Environmental Data Portal

en: GEM2: Meteorological and snow station at Gemsstock (3021 m asl), Canton Uri, Switzerland


maintainer: Nander Wever

en: longwave-radiation air-temperature shortwave-radiation precipitation-amount humidity wind-speed snow-depth wind-direction snow-ice-temperature

en: SNOWPACK sno file

en: This sno-file can be used to run SNOWPACK simulations with the WFJ data set, using version 3.4.0 or higher.

en: SNOWPACK ini file (old syntax)

en: This ini-file can be used to run SNOWPACK simulations with the WFJ data set, using version 3.4.0 or higher.

en: Version 2

en:
Deprecated version. Release notes:

Bug fix: in v1, incoming radiation components were taken from WFJ_Gipfel station, instead of WFJ Versuchsfeld.

en: Version 7 (current)

en:
Release notes:

Bug fix: in previous versions, snow lysimeter data from September 2010 onwards was erroneously reported for UTC+2, instead of UTC+1 (i.e., reported one hour later than should have been).

Update: dataset extended to 2017-07-11T00:30.

en: SNOWPACK ini file (new syntax)

en: This ini-file can be used to run SNOWPACK simulations with the WFJ data set, using the new MeteoIO Filter, Generator and Interpolation syntax.

en: Version 1

en:
Deprecated version. Release notes:

First release of dataset.

en: Version 4

en:
Deprecated version. Release notes:

Update: dataset extended to 2016-09-02T00:30

Update: refreshing data originating from MeteoSwiss (particularly precipitation)

en: Undercatch correction

en: This document describes how the undercatch correction for the Weissfluhjoch measurement site was determined.

en: Version 6

en:
Deprecated version.

Release notes:

- Missing ISWR for 2009-09-30 and 2010-04-30 has been replaced by data from WFJ Gipfel

This dataset is the basis for the Weissfluhjoch dataset in ESM-SnowMIP (doi: http://dx.doi.org/10.16904/16).

en: All versions

en: Version 5

en:
Deprecated version. Release notes:

Update: using the incoming and outgoing short- and longwave radiation data from the new processing chain from 2010 onwards, filling also data gaps in pre-2010 data.

Update: dataset extended to 2016-10-04T00:30

Update: refreshing data originating from MeteoSwiss (particularly precipitation)

en: Version 3

en:
Deprecated version. Release notes:

Bug fix: in version 2, the undercatch correction was not applied correctly

Bug fix: in version 2, snow lysimeter data was missing for extended periods of time

Update: dataset extended to 2016-01-02T00:30

Update: refreshing data originating from MeteoSwiss (particularly precipitation)

en:
Dataset of meteorological and snowpack measurements from the automatic weather station at Weissfluhjoch, Davos, Switzerland, suitable for driving snowpack models. The dataset contains standard meteorological measurements, and additionally snowpack runoff data from a snow lysimeter. Where possible, data is quality checked and missing data are replaced from backup sensors from the measurement site itself, or (in only a few cases) from the MeteoSwiss weather station at 470 m distance and 150 m above the measurement site.

Publication
Wever, N., Schmid, L., Heilig, A., Eisen, O., Fierz, C., and Lehning, M. Verification of the multi-layer SNOWPACK model with different water transport schemes. 2015. The Cryosphere. Volume 9. 2271-2293. http://dx.doi.org/10.5194/tc-9-2271-2015.

air-temperature

display_name: air-temperature

humidity

display_name: humidity

precipitation-amount

display_name: precipitation-amount

snow-depth

display_name: snow-depth

snow-ice-temperature

display_name: snow-ice-temperature

wind-direction

display_name: wind-direction

wind-speed

display_name: wind-speed

en: WFJ_MOD: Meteorological and snowpack measurements from Weissfluhjoch, Davos, Switzerland


en: anthropogenic-management microbial-biomass-carbon nutrient-addition microbial-respiration soil-properties herbivore-exclusion climate

en: Risch et al. GEB_data.xlsx

media_type: .xlsx

en:
Study sites and experimental design
We collected data from 23 sites that are part of the Nutrient Network Global Research Cooperative (NutNet, https://nutnet.umn.edu/). The mean annual air temperature (MAT) across these sites ranged from -4 to 22°C, mean annual precipitation (MAP) from 252 to 1592 mm, and elevations from 6 to 4261 m above sea level (Fig 1a, Supplementary Table S1), hence cover a wide range of climatic conditions under which grasslands occur (Fig 1b). Soil organic C concentrations ranged between 0.8 to 7.8%, soil total N concentrations between 0.1 and 0.6%, and the soil C:N ratio between 9.1 and 21.5. Soil clay content spanned from 3.0 to 35%, and soil pH from 3.4 to 7.6 (Supplementary Table S2).
At each site, the effects of nutrient addition and herbivore exclusion were tested via a randomized-block design (Borer et al., 2014). Three blocks with 10 treatment plots each were established at each site, except for the site at bldr.us (only two blocks). Each of these 10 plots was randomly assigned to a nutrient or fencing treatment. An individual plot was 5 x 5 m, divided into four 2.5 x 2.5 m subplots. Each subplot was further divided into four 1 x 1 m square sampling plots, one of which was set aside for soil sampling (Borer et al., 2014). Plots were separated by at least 1 m wide walkways. We collected soil samples from four different treatments for this study: (i) untreated control plots (Control), (ii) herbivore exclusion plots (Fence), (iii) plots fertilized with N, P, K, plus nine essential macro and micronutrients (NPK), and (iv) plots with simultaneous fertilizer addition and herbivore exclusion (NPK+Fence). The experiments were established at different times in the past, with years of treatment different among sites (2 – 9 years since start of treatment; Supplementary Table S1). For the nutrient additions, all sites applied 10 g N m-2 each year as time-release urea; 10 g P m-2 yr-1 as triple-super phosphate; 10 g K m-2 yr-1 as potassium sulfate. A micro-nutrient mix (Fe, S, Mg, Mn, Cu, Zn, B, Mo, Ca) was applied at 100 g m-2 together with K in the first year of treatments but not thereafter.
We excluded large vertebrate herbivores (Fence) by fencing two plots, one with and one without NPK additions, within each block. The fences excluded all aboveground mammalian herbivores with a body mass of over 50 g (Borer et al., 2014). At most sites, the fences were 180 cm high, and the fence contained a wire mesh (1 cm holes) for the bottom 90 cm with a 30 cm outward-facing flange stapled to the ground to exclude burrowing animals. Climbing and subterranean animals may potentially still access these plots (Borer et al., 2014). For slight modifications in fence design at a few sites see Supplementary Table S3. Most sites only had wild herbivores, although four sites were also grazed by domestic animals (Supplementary Table S1).

Collection of soil samples, soil microbial respiration, microbial biomass, and other soil properties
Each of the 23 sites received a package containing identical material from the Swiss Federal Institute for Forest, Snow and Landscape Research WSL, Switzerland to be used for sampling (Risch et al., 2015; Risch et al., 2019). We collected two soil cores of 5 cm diameter and 12 cm depth in each sampling plot and composited them to measure MR, MBC, and soil chemical properties (see below). An additional sample (5 x 12 cm) was collected to assess soil physical properties. This sample remained within a steel sampling core after collection and both ends were tightly closed with plastic caps to avoid disturbance. All soils were shipped cooled to the laboratory at (Location will be disclosed after manuscript acceptance) within a few days after collection. Soils were sampled roughly 6 weeks prior to peak biomass at each site during 2015 and 2016.
To assess MR (CO2 production) in a laboratory incubation experiment we weighed duplicate soil samples (8 g dry soil equivalent) into 50-ml Falcon tubes. No additional substrate (glucose, sugar) was added to these samples. We adjusted the soil moisture of each sample to 60% field capacity. We then placed a 15 ml plastic test tube (Semadeni 1701A) containing 7.25 ml 0.05 M NaOH over each soil sample. The test tube was fixed with a plastic rod so that it was not in contact with the soil sample. The Falcon tubes were then sealed with a screw cap and placed in an incubator under completely dark conditions at 20°C. The CO2 produced by microbial respiration was absorbed by the 0.05 M NaOH. For five weeks we measured the decrease in conductivity within the 0.05 M NaOH solution on a weekly basis with a Multimeter WTW Multi 3410 (WTW GmbH, Germany) and replaced the 0.05 M NaOH with fresh solution. We included Falcon tubes without soil samples in each incubation run as blanks to test if tubes were tight and no CO2 could enter or escape. We calibrated the relationship between conductivity reduction and NaOH absorbed as follows: 400 ml 0.05 M NaOH was placed in a beaker and its conductivity was measured with the multimeter. While stirring, air containing CO2 was blown into the solution for approximately one minute, which reacts with NaOH to form Na2CO3. After this process, conductivity was measured again. We then transferred 7.25 ml of the solution into a smaller beaker and added 1 ml of 0.1 M BaCl2 to precipitate Na2CO3 and then titrated the solution with 0.05 M HCl to determine the remaining NaOH. We then repeated these steps with the remaining solution a total of nine times and plotted the conductivities (y-axis) against the NaOH consumed (x-axis, Supplementary Fig S1). This regression line was used to infer the consumption of NaOH from the conductivity reduction in the incubation experiments and to calculate CO2 evolution during incubation. In addition, we determined the optimum concentration for the NaOH solution in series of preliminary experiments, so that the concentration was not too high to become insensitive, but also not too low so that not all NaOH reacts during incubation. We then calculated MR (mg CO2-C kg dry soil-1 h-1) as total amount of CO2 released over the 5 weeks divided by the duration of the entire incubation in hrs.
Soil microbial biomass carbon (MBC; mg C kg soil-1 ) was measured at the beginning of the experiment by measuring the maximal respiratory response to the addition of glucose solution (4 mg glucose per g soil dry weight dissolved in distilled water; substrate-induced respiration method) on approximately 5.5 g of soil (J. P. E. Anderson & Domsch, 1978; Nico Eisenhauer et al., 2018; Scheu, 1992). For this purpose we used an O2-micro-compensation apparatus (Scheu, 1992). More specifically, substrate-induced respiration was calculated from the respiratory response to D-glucose for 10 hr at 20°C. Glucose was added according to preliminary studies to saturate the catabolic enzymes of microorganisms (4 mg g soil-1 dissolved in 400 ml deionized water). The mean of the lowest three readings within the first 10 hrs (between the initial peak caused by disturbing the soil and the peak caused by microbial growth) was taken as maximum initial respiratory response (MIRR; ml O2 kg soil-1 h-1) and microbial biomass (mg C kg soil-1) was calculated as 38 x MIRR (Beck et al., 1997; Cesarz et al., 2022; Thakur et al., 2015).
The rest of the composited sample was dried at 65°C for 48 h, ground and sieved (2 mm mesh) to assess the soil pH, mineral soil total C and N and C:N ratio, and mineral soil organic C (Risch et al., 2019). The undisturbed sample was used to assess water holding capacity (WHC), bulk density (BD), and soil texture [sand, silt, clay; methods in (Risch et al., 2019)]. We used the percentage of sand and clay as an indicator of soil texture in this study. MAT (°C), MAP (mm) and temperature of the wettest quarter (°C) were obtained from www.worldclim.com (Fick & Hijmans, 2017; Hijmans, Cameron, Parra, Jones, & Jarvis, 2005). These variables were selected as they were found to be drivers of soil nutrient processes across these sites in earlier studies (Risch et al., 2020; Risch et al., 2019). Mean annual soil temperatures (MAST; °C) for the 0 to 5 cm soil layer were obtained for each site from the SoilTemp maps (J. Lembrechts et al., 2021; J. J. Lembrechts et al., 2022), global gridded modelled products of soil bioclimatic variables for the 1979-2013 period at a 1-km² resolution, based on CHELSA, ERA5 and in-situ soil temperature measurements.
Numerical calculations and statistical analyses
We calculated MMQ as MR/MBC. We corrected this measure using the average soil temperature of each site (MMQsoil). This temperature correction is necessary as incubation temperatures are usually much higher than site mean annual soil temperatures (see Xu et al. 2017). MMQsoil = MMQ x Q10(MAST – 20)/10, where Q10 was assumed to be 2 (Xu et al. 2017). See Supplementary Fig S2 for comparison of air and soil temperatures across the 23 sites as well as the incubation temperature.
Some of the explanatory variables (clay, soil organic C, C:N ratio) were skewed and were thus log-transformed prior to analyses. All continuous explanatory variables were centred and scaled to have a mean of zero and variance of one. To avoid collinearity between them we filtered them using correlation analysis (Supplementary Fig S3). From the variables that were strongly correlated (Pearson’s |r| > 0.70) (Dormann et al., 2013), we selected the ones that allowed us to minimize the number of variables (Supplementary Fig S3). Specifically, soil total N concentration, soil total C concentration, soil sand content and soil bulk density were dropped from the dataset. We then assessed how these edaphoclimatic variables are related to MMQ across our global grasslands.
For this, we used linear mixed effects models (LMMs) fitted by maximum likelihood with the lme function in the nlme package (version 3.1-153) (Pinheiro, Bates, DebRoy, & Sarkar, 2021) in R version 3.6.3. (R Core Team, 2019). We used treatment as a fixed effect and plot nested in site as random effects to assess treatment differences in MMQsoil, as well as MR, and MBC. The number of years since the treatment started was included as a fixed effect in all the initial models but was not significant and therefore not retained in the models. To assess how differences in MMQsoil were affected by environmental factors (soil, climatic properties) we again used LMMs. Soil and climatic properties were included as fixed effects and plot nested in site as random effects. We did not include interactions between environmental variables. We then used the MuMin package (Barton, 2018) (version 1.42.1) to select the best models that explained the most variation based on Akaike’s information criterion (AIC; model.avg function). We used the corrected AIC (AICc) to account for our small sample size and selected the top models that fell within 4 AICc units (delta AICc < 4) (Burnham & Anderson, 2002; Johnson & Omland, 2004). We present all our top models rather than model averages. Conditional averages are provided in the Supplementary material.
Based on findings from analyses described above and the literature, we developed a conceptual model of direct and indirect relationships between both edaphoclimatic variables and experimental treatments (Supplement Figure S4) to obtain a more holistic approach in understanding how these properties affect MMQsoil. We had data from 23 sites with 272 observations. We tested this model using structural equation modelling based on a d-sep approach (Lefcheck, 2016; Shipley, 2009). We considered those environmental drivers that were included in our top LMMs, namely temperature of the wettest quarter (T.q.wet), soil pH, water holding capacity (WHC) and soil organic C (organic C; Supplementary Figure S4). These factors were allowed to directly affect MMQsoil, and via their interactions with treatments. In addition, treatments were allowed to directly affect MMQsoil. Treatments were included as dummy variables in the model. We tested our conceptual model (Supplementary Fig S4) using the piecewiseSEM package (version 2.0.2; Lefcheck, 2016) in R 3.4.0, in which a structured set of linear models are fitted individually. This approach allowed us to account for the nested experimental design, and overcome some of the limitations of standard structural equation models, such as small sample sizes (Lefcheck, 2016; Shipley, 2009). We used the lme function of the nlme package to model response variables, including site as a random factor. Good fit of the SEM was assumed when Fisher’s C values were non-significant (p > 0.05). For all significant interactions between soil or climate variables and treatments detected in the SEMs, we calculated treatment effect sizes, i.e., the differences in MMQsoil between Control and treatments as log response ratios (LRR) and plotted these values against the climate or soil factors. The LRR were defined as log(Control/Treatment), where treatment was either Fence, NPK or NPK+Fence. To assess which of the LRR-climate or soil property relationships were significant we again used LMMs, in which soil and climatic properties were included as fixed effects and plot nested in site as random effects.

en:
This dataset contains all data on which the following publication below is based.

Paper Citation:

Risch Anita C., Zimmermann, Stefan, Schütz, Martin, Borer, Elizabeth T., Broadbent, Arthur A.D., Caldeira, Maria C., Davies, Kendi F., Eisenhauer, Nico, Eskelinen, Anu, Fay, Philip A., Hagedorn, Frank, Knops, Johannes M.H., Lembrechts, Jonas, J., MacDougall, Andrew S., McCulley, Rebecca L., Melbourne, Brett A., Moore, Joslin L., Power, Sally A., Seabloom, Eric W., Silveira, Maria L., Virtanen, Risto, Yahdjian, Laura, Ochoa-Hueso, Raul (accepted). Drivers of the microbial metabolic quotient across global grasslands. Global Ecology and Biogeography

Please cite this paper together with the citation for the datafile.

The microbial metabolic quotient (MMQ; mg CO2-C mg MBC-1 h-1), defined as the amount of microbial CO2 respired (MR; mg CO2-C kg soil-1 h-1) per unit of microbial biomass C (MBC; mg C kg soil-1), is a key parameter for understanding the microbial regulation of the carbon (C) cycle, including soil C sequestration. Here, we experimentally tested hypotheses about the individual and interactive effects of multiple nutrient addition (NPK+micronutrients) and herbivore exclusion on MR, MBC, and MMQ across 23 sites (5 continents). Our sites encompassed a wide range of edaphoclimatic conditions, thus we assessed which edaphoclimatic variables affected MMQ the most and how they interacted with our treatments. Soils were collected in plots with established experimental treatments. MR was assessed in a five-week laboratory incubation without glucose addition, MBC via substrate-induced respiration. MMQ was calculated as MR/MBC and corrected for soil temperatures (MMQsoil). Using LMMs and SEMs, we analysed how edaphoclimatic characteristics and treatments interactively affected MMQsoil. MMQsoil was higher in locations with higher mean annual temperature, lower water holding capacity, and soil organic C concentration, but did not respond to our treatments across sites as neither MR nor MBC changed. We attributed this relative homeostasis to our treatments to the modulating influence of edaphoclimatic variables. For example, herbivore exclusion, regardless of fertilization, led to greater MMQsoil only at sites with lower soil organic C (<1.7%). Our results pinpoint the main variables related to MMQsoil across grasslands and emphasize the importance of the local edaphoclimatic conditions in controlling the response of the C cycle to anthropogenic stressors. By testing hypotheses about MMQsoil across global edaphoclimatic gradients, this work also helps to align the conflicting results of prior studies.

anthropogenic-management

display_name: anthropogenic-management

climate

display_name: climate

herbivore-exclusion

display_name: herbivore-exclusion

microbial-biomass-carbon

display_name: microbial-biomass-carbon

microbial-respiration

display_name: microbial-respiration

nutrient-addition

display_name: nutrient-addition

soil-properties

display_name: soil-properties

education

en: Education and science

envidat

en: EnviDat: the Environmental Data Portal

en: EnviDat is the WSL data portal providing unified and managed access to environmental monitoring and research data. The portal has the capability to host and publish data sets. Through the sharing of unique and high-value data sets worldwide, EnviDat fosters research and contributes to advancing environmental science.


maintainer: Leonie Schönbeck

en: drought pinus-sylvestris photosynthesis vpd fluorescence acclimation gas-exchange

en: Methods ACi curves

en:
Parameters and methods for CO2 response curves of Photosynthesis.

Metatdata format by
Ely et al (2021). A reporting format for leaf-level gas exchange data and metadata. Ecological Informatics. https://doi.org/10.1016/j.ecoinf.2021.101232

en: Methods light response

en: Methods and parameters for light response curves

en: Methods survey

media_type: .csv

en: Parameters and methods for point measurements / survey measurements

en: Instrument details

en:
Instrument details for all LiCor instruments used during collection of dataset.

Metatdata created following guidelines from
Ely et al (2021). A reporting format for leaf-level gas exchange data and metadata. Ecological Informatics. https://doi.org/10.1016/j.ecoinf.2021.101232

en: Variable names and units

media_type: XLSX

en: Variable names, description and units for the Pfynwald Gas exchange data

en: Methods diurnal measurements

en:
Parameters and methodology for diurnal measurements

Metadata format from
Ely et al (2021). A reporting format for leaf-level gas exchange data and metadata. Ecological Informatics. https://doi.org/10.1016/j.ecoinf.2021.101232

en: Pfynwald gas exchange

en:
Full dataset of Pfynwald gas exchange data taken in the period from June 2013 to October 2020.
All metdata can be found in attached documents.
Data uploaded after quality control using guidelines by Ely et al (2021). A reporting format for leaf-level gas exchange data and metadata. Ecological Informatics. https://doi.org/10.1016/j.ecoinf.2021.101232

en: Methods fluorescence

en:
Parameters and methodology for Fluorescence measurements

Metadata format from
Ely et al (2021). A reporting format for leaf-level gas exchange data and metadata. Ecological Informatics. https://doi.org/10.1016/j.ecoinf.2021.101232

en:
Gas exchange was measured on control, irrigated and irrigation-stop trees at the irrigation experiment Pfynwald, during the years 2013, 2014, 2016-2020. The measurement campaigns served different purposes, resulting in a large dataset containing survey data, CO2 response curves of photosynthesis, light response curves of photosynthesis, and fluorescence measurements. Measurements were done with LiCor 6400 and LiCor 6800 instruments. Until 2016, measurements were done on excised branches or branches lower in the canopy. From 2016 onwards, measurements were done in the top of the canopy using fixed installed scaffolds.
All metadata can be found in the attached documents.

acclimation

display_name: acclimation

drought

display_name: drought

fluorescence

display_name: fluorescence

gas-exchange

display_name: gas-exchange

photosynthesis

display_name: photosynthesis

pinus-sylvestris

display_name: pinus-sylvestris

vpd

display_name: vpd

en: 2013-2020 gas exchange at Pfynwald


en: sentinel-2 landsat mowing land-use grassland remote-sensing management use-intensity

en: Preview number of grassland management events 2020

media_type: PNG

en: Spatial distribution of the number of management events for Switzerland and for three zoomed-in regions (a, b, c). The three zoomed-in regions show different grassland-use intensities, which are reflected in the distribution of management frequency (Weber et al., 2023; IN REVIEW)

format: PNG

en: Preview time of 1st grassland management event 2020

en: Spatial distribution of the timing (day of year; DOY) of the first management event for Switzerland and for three zoomed-in regions (a, b, c). The three zoomed-in regions show different grassland-use intensities, which are reflected in the timing of the first event of the year (Weber et al., 2023; IN REVIEW)

en: Grassland-use intensity map 2018

media_type: geotiff

en: Raster stack with information on the number and timing of grassland-management events (i.e. mowing, grazing) at a spatial resolution of 10 m × 10 m for the whole of Switzerland for 2018. See readme file for more details.

format: GeoTIFF

en: Grassland-use intensity map 2021

en: Raster stack with information on the number and timing of grassland-management events (i.e. mowing, grazing) at a spatial resolution of 10 m × 10 m for the whole of Switzerland for 2021. See readme file for more details.

en: Grassland-use intensity map 2019

en: Raster stack with information on the number and timing of grassland-management events (i.e. mowing, grazing) at a spatial resolution of 10 m × 10 m for the whole of Switzerland for 2019. See readme file for more details.

en: Grassland-use intensity map 2020

en: Raster stack with information on the number and timing of grassland-management events (i.e. mowing, grazing) at a spatial resolution of 10 m × 10 m for the whole of Switzerland for 2020. See readme file for more details.

en: Readme

en: Metadata and description of the individual raster layers

en:
A rule-based algorithm [(Schwieder et al., 2022)](https://doi.org/10.1016/j.rse.2021.112795) was used to produce annual maps for 2018–2021 of grassland-management events, i.e. mowing and/or grazing, for Switzerland using Sentinel-2 and Landsat 8 satellite time series. All satellite images were processed with the [FORCE](https://force-eo.readthedocs.io) framework.

The resulting maps provide information on the number and timing of grassland-management events at a spatial resolution of 10 m × 10 m for the whole of Switzerland. For the final maps, permanent grasslands were masked using a variety of land-use layers, according to [Huber et al. (2022)](https://doi.org/10.1002/rse2.298) but replacing the crop mask with the agricultural-use data from the cantons.

We assessed the detection of management events based on independent reference data, which we acquired from daily time series of publicly available webcams that are widely distributed across Switzerland. We further tested the ecological relevance of the generated intensity measures in relation to nationwide biodiversity data (see Weber et al., 2023; IN REVIEW).

grassland

display_name: grassland

land-use

display_name: land-use

landsat

display_name: landsat

management

display_name: management

mowing

display_name: mowing

remote-sensing

display_name: remote-sensing

use-intensity

display_name: use-intensity

education

en: Education and science

envidat

en: EnviDat: the Environmental Data Portal

en: EnviDat is the WSL data portal providing unified and managed access to environmental monitoring and research data. The portal has the capability to host and publish data sets. Through the sharing of unique and high-value data sets worldwide, EnviDat fosters research and contributes to advancing environmental science.

en: Grassland-use intensity maps for Switzerland


global-cryosphere-watch-data-survey

maintainer: Mathias Bavay

title_for_slug: global-cryosphere-watch-data-survey

en: data-usage data-exchange data-portal gcw automatic-publication

en: Third party data usage

en: Data interoperability survey

en: Two surveys on the topic of data usage where conducted for the Global Cryosphere Watch data portal. The first one focused on the data provider point of view while the second one focused on the data user point of view. 37 data providers (ie institutions) worldwide provided their answers for the first survey (from fall 2017 until summer 2018) while 54 users (contacted through various mailing list such as the Cryolist) answered the questions on their third party data usage (fall 2019 until January 2020).

automatic-publication

display_name: automatic-publication

data-exchange

display_name: data-exchange

data-portal

display_name: data-portal

data-usage

display_name: data-usage

gcw

display_name: gcw

en: Global Cryosphere Watch data survey


en: vapor-transport python snow numerical-model pattern-formation

media_type: ZIP

en:
Data and code v1.0 downloaded from the git.

Running the code requires an installation of FEniCS via docker.

en:
This python code uses the Finite Element library FEniCS (via docker) to solve the one dimensional partial differential equations for heat and mass transfer in snow. The results are written in vtk format.

The dataset contains the code and the output data to reproduce the key Figure 5 from the related publication:

_Schürholt, K., Kowalski, J., Löwe, H.; Elements of future snowpack modeling - Part 1: A physical
instability arising from the non-linear coupling of transport and phase changes, The Cryosphere, 2022_

The code and potential updates can be accessed directly through git via:

https://gitlabext.wsl.ch/snow-physics/snowmodel_fenics

numerical-model

display_name: numerical-model

pattern-formation

display_name: pattern-formation

python

display_name: python

snow

display_name: snow

vapor-transport

display_name: vapor-transport

en: A numerical solver for heat and mass transport in snow based on FEniCS