maintainer: Anita Risch

en: swiss-national-park ecosystem-coupling grazing vertebrates invertebrates exclusion subalpine ecosystem-multifunctionality grassland ecosystem-functions

en: Datafile

en:
ID: Plot ID composited of Fence & Treatment number (90 plots total)

Grassland: Name of the six subalpine grasslands where the exclosure setups were erected. In large grasslands (GRIM, MING, STAB), we established four hierarchical exclosure setups (two in short- and two in tall-grass vegetation), whereas in the small grasslands (BOTS, DADA, MARG) only one setup was erected per vegetation type. BOTS = Val dal Botsch, GRIM = Grimmels, MING = Val Mingér, STAB = Alp Stabelchod, DADA = Stabelchod da daint, MARG = Margunet

Fence: Number of exclosure setup (1 - 18). Note that uneven numbers (1,3,5,7,9,11,13,15,17) represent setups established on short-grass vegetation, even numbers (2,4,6,8,10,12,14,16,18) setups established in tall-grass vegetation
Vegetation	Vegetation type, 1 = short-grass vegetation, 2 = tall-grass vegetation; (9 replicate fences on each vegetation type)
Treatment	Progressive animal exclusion, numbers indicate which animals had access to the plot: 1 = Large/medium/small mammals, and invertebrates have access ("L/M/S/I"), 2 = Medium/small mammals, and invertebrates have access (“M/S/I”), 3 = Small mammals and invertebrates have access (“S/I”), 4 = Invertebrates have access (“I”), 5 = No animals have access (“None”)

Bulkden:	Fine fraction soil bulk density (g cm-3)

Soil.moist	: Soil moisture (%)

Soil.org.C: Rhizosphere soil C content (%)

pH: Soil pH

Nema.PC1: First principal component (PC1) of principal component analysis (PCA) of the soil nematode community composition

Nema.PC2: Second principal component (PC2) of principal component analysis (PCA) of the soil nematode community composition

Micro.PC1: 	First principal component (PC1) of principal component analysis (PCA) of the soil microbial community composition

Micro.PC2: Second principal component (PC2) of principal component analysis (PCA) of the soil microbial community composition

Abv.Inv.PC1: First principal component (PC1) of principal component analysis (PCA) of the aboveground-dwelling invertebrate community composition

Abv.Inv.PC2: Second principal component (PC2) of principal component analysis (PCA) of the aboveground-dwelling invertebrate community composition

Arthr.PC1: First principal component (PC1) of principal component analysis (PCA) of the soil arthropod community composition

Arthr.PC2: Second principal component (PC2) of principal component analysis (PCA) of the soil arthropod community composition

Plant.PC1: First principal component (PC1) of principal component analysis (PCA) of the plant community composition

Plant.PC2: Second principal component (PC2) of principal component analysis (PCA) of the plant community composition

Root.biomass: Root biomass (g m-2)

Micro.biomass	: Microbial biomass carbon (mg kg-1)

Plant.richness: Plant species richness

Plant.N: Plant tissue nitrogen content (%)

Soil.N.min: Soil net nitrogen mineralisation (kg ha-1 season)

Soil.resp: Soil carbon dioxide emissions (soil respiration; g m-2 h-1)

en:
This dataset contains all data on which the following publication below is based.

Paper Citation:
 > Risch AC, Ochoa-Hueso R, van der Putten WH, Bump JK, Busse MD, Frey B, Gwiazdowicz DJ, Page-Dumroese DS, Vandegehuchte ML, Zimmermann S, Schütz M. Size-dependent loss of aboveground animals differentially affects grassland ecosystem coupling and functions. 2018. Nature Communications 9: 3684. [doi: 10.1038/s41467-018-06105-4](https://doi.org/10.1038/s41467-018-06105-4).

Please cite this paper together with the citation for the datafile.

#Methods
Study sites
The experimental exclosure setups were installed within the SNP (IUCN category Ia preserve; Dudley 2008), in south-eastern Switzerland. The park covers 172 km2 of forests and subalpine and alpine grasslands along with scattered rock outcrops and scree slopes. The entire area has been protected from human impact (no hunting, fishing, camping or off-trail hiking) since 1914. Large, fairly homogenous patches of short- and tall-grass vegetation, which originate from different historical management and grazing regimes, cover the park’s subalpine grasslands entirely. Short-grass vegetation developed in areas where cattle used to rest (nutrient input) prior to the park’s foundation (14th century to 1914) (Schütz and others 2003, 2006) and is dominated by lawn grass species such as Festuca rubra L., Briza media L. and Agrostis capillaris L. (Schütz and others 2003, 2006). Today, this vegetation type is intensively grazed by diverse vertebrate and invertebrate communities that inhabit the park and consume up to 60% of the available biomass (Risch and others 2013). Tall-grass vegetation developed where cattle formerly grazed, but did not rest, and is dominated by rather nutrient-poor tussocks of Carex sempervirens Vill. and Nardus stricta L. (Schütz and others 2003, 2006). This vegetation type receives considerably less grazing, with only roughly 20% of the biomass consumed (Risch and others 2013). Consequently, the two vegetation types together represent a long-term trajectory of changes in grazing regimes. Underlying bedrock of all grasslands is dolomite, which renders these grasslands rather poor in nutrients regardless of former and current land-use regimes.
Experimental design
To progressively exclude aboveground vertebrate and invertebrate animals, we established 18 size-selective exclosure setups (nine in short-grass, nine in tall-grass vegetation) distributed over six subalpine grasslands across the SNP (Risch and others 2013, 2015). Elevation differences of exclosure locations did not exceed 350 m (between 1975 and 2300 m a.s.l.). The exclosures were established immediately after snowmelt in spring 2009 and were left in place for five consecutive growing seasons (until end of 2013). They were, however, temporarily dismantled every fall (late October after first snowfall) to protect them from avalanches. They were re-established in the same location every spring immediately after snowmelt. Each size-selective exclosure setup consisted of five plots (2 x 3 m) that progressively excluded aboveground vertebrates and invertebrates from large to small. The plots are labelled according to the guilds that had access to them “L/M/S/I”, “M/S/I”, “S/I”, “I”, “None”; L = large mammals, M = medium mammals, S = small mammals, I = invertebrates, None = no animals had access. As we only had permission to have the experimental setup in place for five consecutive growing seasons, the experiment had to be completely dismantled in the late fall of 2013 and all material removed from the SNP.
Our exclosure design was aimed at excluding mammalian herbivores, but naturally also excluded the few medium and small mammalian predators, as well as the entire aboveground invertebrate food web. A total of 26 large to small mammal species can be found in the SNP, but large apex predators are missing (wolf, bear, lynx) . Reptiles, amphibians and birds are scarce to absent in the subalpine grasslands under study. Only two reptile species occur in the park and they are confined to rocky areas that warm up enough for them to survive. One frog species spawns in an isolated pond far from our grasslands. Only three bird species occasionally feed on the subalpine grasslands. Using game cameras (Moultrie 6MP Game Spy I-60 Infrared Digital Game Camera, Moultrie Feeders, Alabaster, AL, USA), we did observe that the medium- and small-sized mammals (marmot/hares and mice) were not afraid to enter the fences and feed on their designated plots. We never spotted reptiles, amphibians or birds on camera. We distinguished between 59 higher aboveground-dwelling invertebrate taxa that our size-selective exclosures excluded (see also methods for aboveground-dwelling invertebrates below).
The “L/M/S/I” plot (not fenced) was located at least 5 m from the 2.1 m tall and 7 x 9 m large main electrical fence that enclosed the other four plots. The bottom wire of this fence was mounted at 0.5 m height and was not electrified to enable safe access for medium and small mammals, while fencing out the large ones. Within each main fence, we randomly established four 2 x 3 m plots separated by 1-m wide walkways from one another and from the main fence line: 1) the “M/S/I” plots were unfenced, allowing access to all but the large mammals; 2) the “S/I” plots (10 x 10 cm electrical mesh fence) excluded all medium-sized mammals. Note that the bottom 10 cm of this fence remained non-electrified to enable safe access for small mammals; 3) the “I” plots (2 x 2 cm metal mesh fence) excluded all mammals. We double-folded the mesh at the bottom 50 cm to reduce the mesh size to smaller than 1 x 1 cm openings; and 4) the “None” plots were surrounded by a 1 m tall mosquito net (1.5 x 2 mm) to exclude all animals. The top of the plot was covered with a mosquito-meshed wooden frame mounted to the corner posts (roof). We treated these plots a few times with biocompatible insecticide (Clean kill original, Eco Belle GmbH, Waldshut-Tiengen, Germany) to remove insects that might have entered during data collection or that hatched from the soil, but amounts were negligible and did not impact soil moisture conditions within these plots.
To assess whether the design of the “None” exclosure (mesh and roof) affected the response variables within the plots and, therefore, influenced the results, we established an additional six “micro-climate control” exclosures (one in each of the six grasslands) (Risch and others 2013, 2015). These exclosures were built as the “None” exclosures but were open at the bottom (20 cm) of the 3-m side of the fence facing away from the prevailing wind direction to allow invertebrates to enter. A 20-cm high and 3-m long strip of metal mesh was used to block access to small mammals. Thus, this construction allowed a comparable micro-climate to the “None” plots, but also a comparable feeding pressure by invertebrates to the “I” plots. We compared various properties within these exclosures against one another to assess if our construction altered the conditions in the “None” plots. We showed that differences in plant (e.g., vegetation height, aboveground biomass) and soil properties (e.g., soil temperature, moisture) found between the “I” and the “None” treatments were not due to the construction of the “None” exclosure, but a function of animal exclusions, although the amount of UV light reaching the plant canopy was significantly reduced (Risch and others 2013).
Aboveground invertebrate sampling
Aboveground invertebrates were sampled with two different methods to capture both ground- and plant-dwelling organisms: 1) we randomly placed two pitfall traps (67 mm in diameter, covered with a roof) filled with 20% propylene glycol in one 1 x 1 m subplot of the 2 x 3 m treatment plots in spring 2013 (May) and emptied them every two weeks until late September 2013 (Vandegehuchte and others 2017b, 2018). A pitfall trap consisted of a plastic cylinder (13 cm depth, 6.75 cm diameter). Within each cylinder we placed a 100 ml plastic vial with outer diameter 6.70 cm and on top of the cylinder we placed a plastic funnel to guide the invertebrates into the vials. Each trap was cover with a cone-shaped and transparent plastic roof to protect the trap from rain (Vandegehuchte and others 2017b, 2018). Note that in the “None” plots only one trap was placed as control to check for effectiveness of the exclosure. 2) We vacuumed all invertebrates from a 60 x 60 cm area on another 1 x 1 m subplot with a suction sampler (Vortis, Burkhard manufacturing CO, Ltd., Rickmansworth, Hertfordshire, UK) every month from June to September 2013 (Vandegehuchte and others 2017b, 2018). For this purpose, we quickly placed a square plastic frame (60 x 60 x 40 cm) with a closable mosquito mesh sleeve attached to the top edge into the plot from the outside. The suction sample was then inserted into through the sleeve and operated for 45 s to collect the invertebrates (Vandegehuchte and others 2017b, 2018).
We sorted the ≈100 000 individuals collected with both methods by hand and identified each individual morphologically to the lowest taxonomic level feasible (59 taxa, including orders, suborders, subfamilies, families; phylum for Mollusca). These taxa belonged to the following feeding types: 19 herbivores, 16 detritivores, 9 predators, 8 mixed feeders, 5 omnivores and 2 non-classified feeders (or not feeding as adults) (Vandegehuchte and others 2017b). We summed the numbers from the two pitfall traps and the suction sampling over the course of the 2013 season to represent the aboveground invertebrate abundance and community composition of a plot. Note: we did not specifically attempt to catch flying invertebrates with e.g., sticky traps, thus a few flying insects may have been missed with our vacuum sampling approach.
Sampling of plant properties
The vascular plant species composition was assessed at peak biomass every summer (July) by estimating the frequency of occurrence of each species with the pin count method in each plot (Frank and McNaughton 1990). A total of 172 taxa occurred within our 90 plots and we calculated plant species richness for each plot separately. We used the 2013 data in this study. Plant quality was assessed every year in July and September; here we use plant quality at the end of the experiment (September 2013). Two 10 x 100 cm wide strips of vegetation per plot were clipped, combined, dried at 65°C, and ground (Pulverisette 16, Fritsch, Idar-Oberstein, Germany) to pass through a 0.5 mm sieve. Twenty randomly selected samples across all treatments were analysed for N (Leco TruSpec Analyser, Leco, St. Joseph, Michigan, USA) (Vandegehuchte and others 2015). Nitrogen concentrations of the other samples were then estimated from models established for the experiment and the entire SNP relating Fourier transform-near infrared reflectance (FT-NIR) spectra to the measured values of N using a multi-purpose FT-NIR spectrometer (Bruker Optics, Fällanden, Switzerland) (Vandegehuchte and others 2015). Root biomass was sampled every fall by collecting five 2.2 cm diameter x 10 cm deep soil samples (Giddings Machine Company, Windsor, CO, USA) per plot (450 samples year-1). The samples were dried at 30 °C and roots were sorted from the sample by hand. We sorted each sample for 1 h which allowed to retrieve over 90% of all roots present in the samples (Risch and others 2013). The roots were then dried at 65 °C for 48 and weighed to the nearest mg. We averaged the values per plot and used the 2013 data only in this study.
Sampling of edaphic communities
In 2009, 2010, and 2011 we collected three composited soil samples (5 cm diameter x 10 cm depth; AMS Samplers, American Falls, ID, USA) and assessed bacterial community structure using T-RFLP profiling (Liu and others 1997; Blackwood and others 2003; Hodel and others 2014). We detected a total of 89 operational taxonomic units (OTUs). These values are in accordance with other studies reporting OTU richness (Wirthner and others 2011; Zumsteg and others 2012; Meola and others 2014) using T-RFLP profiling, a method that detects the most abundant, and thus likely, the most relevant, taxa. We averaged the data over the three years of collections for our calculations. Microbial biomass carbon (MBC) was determined with the substrate-induced method (Anderson and Domsch 1978) every fall (September) between 2009 and 2013 by collecting three mineral soil samples (5 cm diameter × 10 cm mineral soil core, AMS Samplers, American Falls, ID, USA). The three samples were combined (90 samples for each sampling year), immediately put on ice, taken to the laboratory, passed through a 2-mm sieve and stored at 4°C. Again, we only used the 2013 data in this study.
Soil samples (5 cm diameter x 10 cm depth) to extract soil arthropods were collected in June, July, and August 2011 with a soil corer lined with a plastic sleeve to ensure an undisturbed sample (total of 270 samples). The plastic line core was immediately sealed on both ends using cling film and put into a cooler. All plots were sampled within three days and the extraction of arthropods started the evening of the sampling day using a high-gradient Tullgren funnel apparatus (Crossley and Blair 1991; Vandegehuchte and others 2015). Samples were kept in the extractor for four days and the soil arthropods were collected in 95% ethanol. All individuals were counted and each individual was identified morphologically to the lowest level feasible [76 taxa, including orders, suborders, subfamilies, families (Protura, Thysanoptera, Aphidina, Psylina, Coleoptera, Brachycera, Nematocera, Auchenorryncha, Heteroptera, Formicidae); sub-phylum for Myriapoda, for Acari and Collembola also including morpho-species). Note that we also included larval stages (nine of the 76 taxa) (Vandegehuchte and others 2015). All data were summed over the season. A detailed species list for mites and collembolans is published (Vandegehuchte and others 2017a) [https://doi.org/10.1371/journal.pone.0118679.s001]. Earthworms are rare in the SNP and therefore were not included. We collected eight random 2.2 cm diameter x 10 cm deep soil cores from each plot in September 2013 to determine the soil nematode community composition. The samples were mixed and the nematodes were extracted from 100 ml of fresh soil using Oostenbrink elutriators (Oostenbrink 1960). All nematodes in a 1 ml of the 10 ml extract were counted, a minimum of 150 individuals sample-1 were identified to genus or family level using (Bongers 1988), the numbers of all nematodes were extrapolated to the entire sample and expressed for a 100 g dry sample. In total we identified 63 genus or family levels (Vandegehuchte and others 2015). The list of all the nematodes found is published (Vandegehuchte and others 2015) [http://www.oikosjournal.org/appendix/oik-03341] or DOI: [doi: 10.1111/oik.03341].
We are aware that sampling soil microbes from 2009 to 2011 and soil arthropods in 2011 was not ideal, but we are positive that this does not bias the results. Most of the parameters measured in our experiment either already showed a treatment response after the first growing season (e.g., plant biomass) or did not respond over the entire time experiment (e.g., microbial biomass C). The microbial community composition (2009 – 2011) was highly influenced by inter-annual differences in temperature and precipitation, but did not differ between treatments or vegetation types (Hodel and others 2014). We therefore felt comfortable using the 2009 through 2011 data for describing the soil microbial community in our experimental treatments. Similarly, we are positive that our soil arthropod data are representative. We did assess soil arthropods in August 2012 and found no differences to the August 2011 data. However, we did not feel comfortable combining the 2011 June, July, August data with only August data for 2012 for our analyses.
Sampling of soil properties
We collected three soil samples (5 cm diameter x 10 cm depth) in each plot in September 2013 after removing the vegetation. First, we collected the top layer of mineral soil rich in organic matter, the surface organic layer or rhizosphere, typically 1 to 3 cm in depth with a soil corer (AMS Samples, American Falls, Idaho, USA). Second, we collected a 10 cm mineral soil core beneath this surface layer. The cores for each layer were composited, dried at 65 °C for 48 h and fine-ground to pass a 0.5 mm screen. We then analysed all samples for total C using a Leco TruSpec Analyser (Leco, St. Joseph, Michigan, USA). Mineral soil pH was measured potentiometrically in 1:2 soil:CaCl2 solution with an equilibration time of 30 min.
Soil net N mineralisation was assessed during the 2013 growing season (Risch and others 2015). For this purpose, we randomly collected a 5 cm diameter x10 cm deep soil sample with a soil corer (AMS Samples, American Falls, Idaho, USA) after clipping the vegetation in June 2013. After weighing and sieving (4 mm mesh) the soil, we extracted a 20 g subsample in 1 mol l-1 KCl for 1.5 h on an end-over-end shaker and thereafter filtered it through ashless folded filter paper (DF 5895 150, ALBET LabScience, Hahnenmühle FineArt GmbH, Dassel, Germany). From these filtrates NO3- concentrations were measured colorimetrically (Norman and Stucki 1981) and NH4+with flow injection analysis (FIAS 300, Perkin Elmer, Waltham Massachusetts, USA) (Risch and others 2015). We dried the rest of the sample 105 °C to constant mass to determine fine,fraction bulk density. A second soil sample was collected within each plot in June 2013 with a corer lined with a 5 x 13 cm aluminium cylinder. The corer was driven 11.5 cm deep into the soil so that the top 1.5 cm of the cylinder remained empty. Into this space we placed a polyester bag (250 µm) filled an ion-exchanger resin to capture the incoming N. The bag was filled with a 1:1 mixture of acidic and alkaline exchanger resin (ion-exchanger I KA/ion exchanger IIIAA, Merck AG, Darmstadt, Germany). We then removed 1.5 cm soil at the bottom of the cylinder and placed a second resin exchanger bag into this space to capture the N leached from the soil column. To assure that the exchange resin was saturated with H+ and Cl- prior to filling the bags, the mixture was stirred with 1.2 ml l-1 HCl for 1 h and then rinsed with demineralized water until the electrical conductivity of the water reached 5 µm cm-1. The cylinder with the resin bags in place was reinserted into the soil with the top flush to the soil surface and incubated for three months. We recollected the cylinders in September 2013. Each resin bag and 20 g of sieved soil (4 mm mesh) from each cylinder were then separately extracted with KCl and NO3- and NH4+ concentrations were measured. Nitrate and NH4+ concentrations of all samples were then converted to a content basis by multiplying their values with fine fraction bulk density. Net N mineralisation was thereafter calculated as the difference between the N content of the samples collected at the end of the three-month incubation (including the N extracted from the bottom resin bag) and the N content at the beginning of the incubation (Risch and others 2015).
Soil CO2 emissions were measured every two weeks between 0900 and 1700 hrs from early May through late September 2013 with a PP-Systems SRC-1 soil respiration chamber (15 cm high, 10 cm diameter; closed circuit) attached to a PP-Systems EGM-4 infrared gas analyser (PP-Systems, Amesbury, MA, USA) on two locations per plot (Risch and others 2013). The chamber was placed on randomly placed, permanently installed PVC collars (10 cm diameter) driven 5 cm into the soil at the beginning of the study (Risch and others 2013). Freshly germinated plants growing within the collars were removed prior to each measurement to avoid measuring plant respiration or photosynthesis. The two measurements collected per plot and sampling date were averaged.
Soil moisture (with time domain reflectometry; Field-Scout TDR-100, Spectrum Technologies, Plainfield, Illionois, USA) and temperature (with a waterproof digital pocket thermometer; Barnstead International, Dubuque, Iowa, USA) were measured at five random locations per plot every two weeks during the growing seasons during the experiment for the 0 to 10 cm depth (Risch and others 2013, 2015). As soil moisture and soil temperature were highly negatively correlated (Risch and others 2013), we only used soil moisture for this study. We used plot-level averages of all values available to capture soil moisture variability during the five years of the experiment. The results remained unchanged when we only used soil moisture from the 2013 growing season.
Numeral calculations and statistical analyses
Ecosystem coupling. We conducted principal component analyses (PCAs; unscaled) at the complete dataset level using the abundances of each taxonomical entity to describe each of the five different communities used in this study: aboveground-dwelling invertebrates, vascular plants, soil microorganisms, soil arthropods and soil nematodes. We retained the first two components (PCA axis 1 and PCA axis 2) of each analysis as we found them to adequately represent the temporal and spatial variability of our 90 treatment plots in previous studies55,67. Together they explained a total of 71.70% of the variation for aboveground invertebrates, 44.36% for plants, 44.85% for soil microorganisms, 61.85% for soil arthropods and 77.19% for soil nematodes. In addition, we used soil pH and soil organic C content as a proxy for soil chemical properties, soil bulk density as a proxy for soil physical properties and soil moisture (negatively correlated with soil temperature) as a proxy for soil micro-climatic conditions for an overall total of fourteen constituents.
We calculated ecosystem coupling9 for each exclosure treatment within each vegetation type (i.e., 2  5 treatment combinations in total) as an integrated measure of pairwise ecological interactions between ecosystem constituents representing ecological communities and the soil abiotic environment. These ecological interactions are defined by non-parametric Spearman rank correlation analyses between two constituents, excluding interactions involving two abiotic constituents (e.g., soil pH vs. soil moisture) and interactions between the first (PC1) and second (PC2) component of each community type, as these are orthogonal by definition. Interactions between abiotic constituents were excluded from the analyses because the focus of our study was on communities and how they interact with one another and their surrounding environment; therefore, including abiotic-abiotic interactions was not of interest here. Given that the effectiveness of our experimental design resulted in that no community composition data of aboveground-dwelling invertebrates was available for the “None” plots (all animals excluded), only thirteen instead of fourteen constituents were included in the ecosystem coupling calculations for this treatment. The complete absence of aboveground invertebrates represents the most extreme case of disturbance between aboveground animal communities and the rest of the ecosystem constituents. This may have resulted in a slight overestimation of ecosystem coupling for these plots.
	Average ecosystem coupling was calculated as follows:
Ecosystem coupling=
where Xi is the absolute Coupling was calculated value of the Spearman’s rho coefficient of the ith correlation for each treatment within each vegetation type (i.e., based on nine replicates each), considering and n is the number of pairwise comparisons considered (n = a total of 80; interactions (56 in the case of the “None” treatment). We considered a total of 40 biotic-biotic interactions (i.e., concerning two community-level principal components such as plants and microbes; 24 in the case of the “None” treatment) and 40 abiotic-biotic (i.e., concerning one community-level principal component and one abiotic factor, e.g., plant community and soil properties; 32 in the case of the “None” treatment).	Coupling was calculated for each treatment within each vegetation type (i.e., based on nine replicates each), considering a total of 80 interactions (56 in the case of the “None” treatment). We considered a total of 40 biotic-biotic interactions (i.e., concerning two community-level principal components such as plants and microbes; 24 in the case of the “None” treatment) and 40 abiotic-biotic (i.e., concerning one community-level principal component and one abiotic factor, e.g., plant community and soil properties; 32 in the case of the “None” treatment).
To establish whether constituents were significantly and positively coupled within treatments (i.e., the average of their correlation coefficients were greater than in a null model where correlation only happens by chance), we calculated one-tailed p-values based on permutation tests with 999 permutations.

We considered six ecosystem functions and process rates commonly used to assess ecosystem functioning (Meyer and others 2015; Manning and others 2018). Plant N content represents a measure of forage quality, while plant richness has been shown to stabilise biomass production, thus allowing the system to respond to changes in herbivory. Soil net N mineralisation, soil respiration, root biomass, and microbial biomass represent fluxes or stocks of energy. For all functions and processes higher values represent higher functioning (Manning and others 2018). All these variables were measured in the last year of the experiment (2013). We then quantified ecosystem multifunctionality using the multiple threshold approach (Byrnes and others 2014; Manning and others 2018), which considers the number of functions that are above a certain threshold, over a series of threshold values (typically 10-99%) that are defined based on the maximum value of each function. We weighted all our functions equally for these calculations (Manning and others 2018). The number of functions in a plot with values higher than a given threshold value for the respective function is summed up. The sum represents ecosystem multifunctionality for that plot. Given that choosing any particular threshold as a measure of ecosystem multifunctionality is arbitrary, we calculated the average of thresholds from 10-90% (in 10% intervals) as a more integrated representation of ecosystem multifunctionality.
We used Pearson correlations to explore the relationships between ecosystem coupling (all interactions, biotic-biotic interactions, abiotic-biotic interactions involving above- and belowground constituents, and all interactions, biotic-biotic interactions, abiotic-biotic interactions involving belowground constituents only) and ecosystem multifunctionality by calculating the slopes of all relationships between ecosystem coupling and multifunctionality for all thresholds between 10 and 99%. We also related ecosystem coupling with the average of multifunctionality at thresholds between 30-80% as explained before and considered this correlation as a robust indication of the type of association between these two variables. In addition, we explored the relationships between ecosystem coupling (all interactions, biotic-biotic interactions, abiotic-biotic interactions involving above- and belowground constituents, and all interactions, biotic-biotic interactions, abiotic-biotic interactions involving belowground constituents only) and individual ecosystem functions. The effects of exclosures and vegetation type on individual functions and multifunctionality were evaluated using linear mixed effects models ('lme' function of the nlme package), with exclosure and vegetation type as fixed effects and fence as a random factor. All statistical analyses and numerical calculations were done in R version 3.4.0 (R Core Team 2016).

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- Risch AC, Schütz M, Vandegehuchte ML, Van Der Putten WH, Duyts H, Raschein U, Gwiazdowicz DJ, Busse MD, Page-Dumroese DS, Zimmermann S. 2015. Aboveground vertebrate and invertebrate herbivore impact on net N mineralization in subalpine grasslands. Ecology 96:3312–22.
- Schütz M, Risch AC, Achermann G, Thiel-Egenter C, Page-Dumroese DS, Jurgensen MF, Edwards PJ. 2006. Phosphorus translocation by red deer on a subalpine grassland in the Central European Alps. Ecosystems 9:624–633.
- Schütz M, Risch AC, Leuzinger E, Krüsi BO, Achermann G. 2003. Impact of herbivory by red deer (Cervus elaphus L.) on patterns and processes in subalpine grasslands in the Swiss National Park. For Ecol Manage 181:177–88.
- Vandegehuchte ML, van der Putten WH, Duyts H, Schütz M, Risch AC. 2017a. Aboveground mammal and invertebrate exclusions cause consistent changes in soil food webs of two subalpine grassland types, but mechanisms are system-speciﬁc. Oikos 126:212–23.
- Vandegehuchte ML, Raschein U, Schütz M, Gwiazdowicz DJ, Risch AC. 2015. Indirect short- and long-term effects of aboveground invertebrate and vertebrate herbivores on soil microarthropod communities. PLoS One 10:e0118679.
- Vandegehuchte ML, Schütz M, de Schaetzen F, Risch AC. 2017b. Mammal-induced trophic cascades in invertebrate food webs are modulated by grazing intensity in subalpine grassland. J Anim Ecol 86:1434–46.
- Vandegehuchte ML, Trivellone V, Schütz M, Firn J, de Schaetzen F, Risch AC. 2018. Mammalian herbivores affect leafhoppers associated with specific plant functional types at different timescales. Funct Ecol 32:545–55.
- Wirthner S, Frey B, Busse MD, Schütz M, Risch AC. 2011. Effects of wild boar (Sus scrofa L.) rooting on the bacterial community structure in mixed-hardwood forest soils in Switzerland. Eur J Soil Biol 47:296–302. http://dx.doi.org/10.1016/j.ejsobi.2011.07.003
- Zumsteg A, Luster J, Göransson H, Smittenberg RH, Brunner I, Bernasconi SM, Zeyer J, Frey B. 2012. Bacterial, Archaeal and Fungal Succession in the Forefield of a Receding Glacier. Microb Ecol 63:552–64. https://doi.org/10.1007/s00248-011-9991-8

ecosystem-coupling

display_name: ecosystem-coupling

ecosystem-functions

display_name: ecosystem-functions

ecosystem-multifunctionality

display_name: ecosystem-multifunctionality

exclusion

display_name: exclusion

grassland

display_name: grassland

grazing

display_name: grazing

invertebrates

display_name: invertebrates

subalpine

display_name: subalpine

swiss-national-park

display_name: swiss-national-park

vertebrates

display_name: vertebrates

en: Ecosystem coupling and multifunctionality - exclosure experiment


maintainer: Christian Rixen

en: nitrogen-isotope tree-rings dwarf-shrubs stillberg fertilization treeline

state: active

type: dataset

en: Metadata Nutrient Addition Stillberg Experimental Design

media_type: image/png

en: Nutrient addition map of the study area with the investigated plots. Each circle (study block) contains one control plot and two plots that received NPK fertilizer corresponding to 15 (NPK15) or 30 (NPK30) kg N ha-1 a-1. Contour lines specify the elevation (m a.s.l.) in steps of 10 m.

format: PNG

en: Project Description Nutrient Addition Stillberg

media_type: text/html

en: Link to Nutrient Addition project description on the WSL website (in German).

format: HTML

en: Data Nutrient Addition Stillberg

en: This link leads to the figshare repository, which contains all nutrient addition data.

format: SERVICE

en:
 Background information

The availability of nitrogen (N) and phosphorus (P) is considered to be a major factor limiting growth and productivity in terrestrial ecosystems globally. This project aimed to determine whether the growth stimulation documented in previous short‐term fertilisation trials persisted in a longer‐term study (12 years) in the treeline ecotone, and whether possible negative effects of nutrient addition offset the benefits of any growth stimulation.
Over the course of the 12 study years, NPK fertiliser corresponding to 15 or 30 kg N ha−1 a−1 was added annually to plots containing 30‐year‐old *Larix decidua* or 32‐year-old *Pinus uncinata* individuals with an understorey of mainly ericaceous dwarf shrubs. To quantify growth, annual shoot increments of trees and dwarf shrubs as well as radial growth increments of trees were measured. Nutrient concentrations in the soil were also measured and the foliar nutritional status of trees and dwarf shrubs was assessed.

 Experimental design

Over an elevation gradient of 140 m across the treeline afforestation site Stillberg, 22 locations were chosen that covered the whole range of microenvironmental conditions (*see* Nutrient addition experimental design.png). Half of the blocks included European larch (*L. decidua*) and the other half included mountain pine (*P. uncinata*). Within each block, three plantation quadrats were randomly selected as experimental plots and each plot was assigned to a control (no fertilisation) or to one of two fertiliser dose treatments (15 kg and 30 kg N ha−1 a−1). Treatments were assigned randomly but confined so that the location of fertilised plots within a block was not directly above control plots to avoid nutrient input from drainage. For details about the experiment, *see* Möhl et al (2019).

 Data description

The available datasets contain climate variables (2004-2016), nutrient isotope measurements (2010 & 2016), shrub growth measurements (2004-2016), soil parameter measurements and annual ring and shoot measurements (2004-2016). All data can be found here:
<https://figshare.com/articles/dataset/Twelve_years_of_low_nutrient_input_stimulates_growth_of_trees_and_dwarf_shrubs_in_the_treeline_ecotone/7025858>

dwarf-shrubs

display_name: dwarf-shrubs

fertilization

display_name: fertilization

nitrogen-isotope

display_name: nitrogen-isotope

stillberg

display_name: stillberg

tree-rings

display_name: tree-rings

treeline

display_name: treeline

educ

fr: Éducation, culture et sport

de: Bildung, Kultur und Sport

en: Education, culture and sport

it: Istruzione, cultura e sport

envidat

fr: EnviDat: le portail de données environnementales

de: EnviDat: das Umwelt-Datenportal

en: EnviDat: the Environmental Data Portal

it: EnviDat: il portale dei dati ambientali

fr: EnviDat est le portail de données du WSL; il offre un accès uniformisé et réglementé aux données de surveillance et de recherche en matière d’environnement. Ce portail est capable d’héberger et de publier des ensembles de données précieuses et uniques, qui sont partagées aux quatre coins de la planète. Ce faisant, EnviDat favorise la recherche et contribue à l’avancement des sciences environnementales.

en: EnviDat is the WSL data portal providing unified and managed access to environmental monitoring and research data. The portal has the capability to host and publish data sets. Through the sharing of unique and high-value data sets worldwide, EnviDat fosters research and contributes to advancing environmental science.

de: EnviDat ist das Datenportal der WSL für einen einheitlichen und verwalteten Zugang zu Umweltdaten aus Monitoring- und Forschungsprojekten. Auf dem Portal können Datensätze gehostet und veröffentlicht werden. Durch die weltweite Bereitstellung einzigartiger und wertvoller Datensätze leistet EnviDat einen Beitrag zur Forschung im Bereich der Umweltwissenschaften.

it: EnviDat è il portale dati del WSL che fornisce un accesso unificato e gestito ai dati di monitoraggio ambientale e di ricerca. Il portale ha la capacità di ospitare e pubblicare serie di dati preziosi e unici che vengono condivisi in tutto il mondo. Così facendo, EnviDat promuove la ricerca e contribuisce al progresso della scienza ambientale.

political_level: other

wsl

capacity: public

type: organization

approval_status: approved

key: guid

key: uri

value: Envidat Harvester

en: Nutrient addition experiment at the Alpine treeline site Stillberg, Switzerland


maintainer: Athanasios Nenes

en: sigma-w raclets supersaturation cloud-modeling cloud-droplets wolfgangpass

en: Predicted cloud droplet numbers (kappa=0.25) Weissfluhjoch

media_type: text/csv

en:
In-situ measurements of aerosol size distributions along with updraft velocities collected from a wind Doppler Lidar are coupled with a state-of-the art droplet activation parameterization (Morales and Nenes, 2014) to determine the cloud droplet number and maximum in-cloud supersaturation that can potentially form over the Davos Wolfgang station between February 24 and March 8 2019. For this, as a sensitivity test, a hygroscopicity parameter of 0.25 is assumed, which is the upper limit determined from the analysis carried out for Weissfluhjoch (https://www.envidat.ch/dataset/ccn-hygroscopicity-predicted-cloud-droplet-numbers-weissfluhjoch). The data file contains:

* Date and time (DD/MM/YYYY hh:mm:ss) in UTC
* The wind-lidar derived standard deviation of updraft velocities (sigma-w) in m/s, used to apply the parameterization employing a single characteristic velocity (Morales and Nenes, 2010)
* The maximum predicted cloud parcel supersaturation (Smax%)
* The total in-situ aerosol number concentration (Naer) in cmˉ³, measured by the SMPS
* The predicted cloud droplet numbers (Nd) in cmˉ³
* The activation fraction (%)

format: CSV

mimetype: text/csv

en: Predicted cloud droplet numbers (kappa=0.1) Davos Wolfgang

en:
In-situ measurements of aerosol size distributions along with updraft velocities collected from a wind Doppler Lidar are coupled with a state-of-the art droplet activation parameterization (Morales and Nenes, 2014) to determine the cloud droplet number and maximum in-cloud supersaturation that can potentially form over the Davos Wolfgang station between February 24 and March 8 2019. For this, as a sensitivity test, a hygroscopicity parameter of 0.1 is assumed, which is the lower limit determined from the analysis carried out for Weissfluhjoch (https://www.envidat.ch/dataset/ccn-hygroscopicity-predicted-cloud-droplet-numbers-weissfluhjoch). The data file contains:

* Date and time (DD/MM/YYYY hh:mm:ss) in UTC
* The wind-lidar derived standard deviation of updraft velocities (sigma-w) in m/s, used to apply the parameterization employing a single characteristic velocity (Morales and Nenes, 2010)
* The maximum predicted cloud parcel supersaturation (Smax%)
* The total in-situ aerosol number concentration (Naer) in cmˉ³, measured by the SMPS
* The predicted cloud droplet numbers (Nd) in cmˉ³
* The activation fraction (%)

en: Cloud droplet properties were predicted between February 24 and March 8 2019 for the measurement site Davos Wolfgang (1630 m a.s.l., LON: 9.853594, LAT: 46.835577). Droplet calculations are carried out with the physically based aerosol activation parameterization of Morales and Nenes (2014), employing the “characteristic velocity” approach of Morales and Nenes (2010). Aerosol size distribution observations required to predict the cloud droplet numbers and maximum in-cloud supersaturation are obtained from a Scanning Mobility Particle Size Spectrometer (SMPS) instrument deployed at Davos Wolfgang (https://www.envidat.ch/dataset/aerosol-data-davos-wolfgang). The required vertical velocity measurements are derived from the wind Doppler Lidar (https://www.envidat.ch/dataset/lidar-wind-profiler-data) deployed at the same station and are extracted from the first bin of the instrument, being 200 m above ground level. The hygroscopic properties of the particles measured at Davos Wolfgang could not be estimated, owing to a lack of concurrent CCN measurements. As a sensitivity test, droplet calculations are performed assuming two different values of the aerosol hygroscopicity parameter, 0.1 and 0.25, based on the analysis carried out for Weissfluhjoch. Additional information can be found in Section 2.3 [here](https://acp.copernicus.org/preprints/acp-2020-1036/).

cloud-droplets

display_name: cloud-droplets

cloud-modeling

display_name: cloud-modeling

raclets

display_name: raclets

sigma-w

display_name: sigma-w

supersaturation

display_name: supersaturation

wolfgangpass

display_name: wolfgangpass

en: Predicted cloud droplet numbers Davos Wolfgang


maintainer: EnviDat Support

en: switzerland climate-change alpine3d-simulations graubunden snow

en: Configuration file

en: Simulation data

en: Reference data as well as the 2050 and 2099 scenarios for ARPEGE and ETHZ other RCMs available on demand)

en:
This is the simulation dataset from _"Response of snow cover and runoff to climate change in high Alpine catchments of Eastern Switzerland"_, M. Bavay, T. Grünewald, M. Lehning, Advances in Water Resources 55, 4-16, 2013

A model study on the impact of climate change on snow cover and runoff has been conducted for the Swiss Canton of Graubünden. The model Alpine3D has been forced with the data from 35 Automatic Weather Stations in order to investigate snow and runoff dynamics for the current climate. The data set has then been modified to reflect climate change as predicted for the 2021-2050 and 2070-2095 periods from an ensemble of regional climate models.

The predicted changes in snow cover will be moderate for 2021-2050 and become drastic in the second half of the century. Towards the end of the century the snow cover changes will roughly be equivalent to an elevation shift of 800 m. Seasonal snow water equivalents will decrease by one to two thirds and snow seasons will be shortened by five to nine weeks in 2095.

Small, higher elevation catchments will show more winter runoff, earlier spring melt peaks and reduced summer runoff. Where glacierized areas exist, the transitional increase in glacier melt will initially offset losses from snow melt. Larger catchments, which reach lower elevations will show much smaller changes since they are already dominated by summer precipitation.

climate-change

display_name: climate-change

graubunden

display_name: graubunden

snow

display_name: snow

switzerland

display_name: switzerland

en: Alpine3D simulations of future climate scenarios for Graubunden


maintainer: M. Carol Resch

en: nematodes restoration topsoil-removal long-term-experiment grassland vegetation diversity

en: Nemadotes and vegetation

en:
Plot: Plot ID combining treatment code and plot number (in total 55 plots)

Treatment: Three restoration treatments and two reference treatments. Initial=intensively managed grasslands; Harvest only= multiple hay harvest (2-3 times) per years; Topsoil= Topsoil removal (upper 10-30 cm); Topsoil+Propagules= Topsoil removal (upper 10-30 cm) combined with introduction of target plant species; Target= species-rich ancient grasslands

X: X-Coordinate

Y: Y-Coordinate

Ri.veg: Total number of plant species

Div.veg: Shannon diversity of plant species

Ri.nem: Total number of nematode families

Div.nem: Shannon diversity of nematode families

abund: Total nematode abundance (individuals per 100g dry soil)

MI: Maturity index according to Bongers 1990; calculations done in online tool NINJA (Sieriebriennikov, Ferris, & de Goede, 2014)

PPI: Plant parasite index according to Bongers 1990; calculations done in online tool NINJA (Sieriebriennikov, Ferris, & de Goede, 2014)

PPI.MI: Plant parasite to Maturity index according to Bongers, van der Meulen, & Korthals 1997; calculations done in online tool NINJA (Sieriebriennikov, Ferris, & de Goede, 2014)

CI: Channel index according to Ferris, Bongers, & de Goede 2001 (%); calculations done in online tool NINJA (Sieriebriennikov, Ferris, & de Goede, 2014)

EI: Enrichment index according to Ferris, Bongers, & de Goede 2001 (%); calculations done in online tool NINJA (Sieriebriennikov, Ferris, & de Goede, 2014)

SI: Structure index according to Ferris, Bongers, & de Goede 2001 (%); calculations done in online tool NINJA (Sieriebriennikov, Ferris, & de Goede, 2014)

He.a: Herbivorous nematode abundance (individuals per 100g dry soil); classification according to Sieriebriennikov, Ferris, & de Goede, 2014

He.ecto.a: Herbivorous ectoparasite nematode abundance (individuals per 100g dry soil); classification according to Sieriebriennikov, Ferris, & de Goede, 2014

He.endo.a: Herbivorous endoparasite nematode abundance (individuals per 100g dry soil); classification according to Sieriebriennikov, Ferris, & de Goede, 2014

Ba.a: Bacterivorous nematode abundance (individuals per 100g dry soil); classification according to Sieriebriennikov, Ferris, & de Goede, 2014

Fu.a: Fungivorous nematode abundance (individuals per 100g dry soil); classification according to Sieriebriennikov, Ferris, & de Goede, 2014

Om.a: Omnivorous nematode abundance (individuals per 100g dry soil); classification according to Sieriebriennikov, Ferris, & de Goede, 2014

Ca.a: Carnivorous nematode abundance (individuals per 100g dry soil); classification according to Sieriebriennikov, Ferris, & de Goede, 2014

Om.Ca.a	Sum of omnivorous and carnivorous nematode abundance (individuals per 100g dry soil); classification according to Sieriebriennikov, Ferris, & de Goede, 2014
CP1.a: C-P (colonizer-persister) class 1 nematode abundance (individuals per 100g dry soil); C-P classification according to Bongers 1990; equivalent to enrichment indicators according to Ferris, Bongers, & de Goede 2001

CP2.a: C-P (colonizer-persister) class 2 nematode abundance (individuals per 100g dry soil); C-P classification according to Bongers 1990; equivalent to stress indicators according to Ferris, Bongers, & de Goede 2001

CP3.a: C-P (colonizer-persister) class 3 nematode abundance (individuals per 100g dry soil); C-P classification according to Bongers 1990

CP4.a: C-P (colonizer-persister) class 4 nematode abundance (individuals per 100g dry soil); C-P classification according to Bongers 1990

CP5.a: C-P (colonizer-persister) class 5 nematode abundance (individuals per 100g dry soil); C-P classification according to Bongers 1990

CP3-5.a: Sum of C-P3, C-P4 and C-P5 nematode abundance (individuals per 100g dry soil) equivalent to structure indicators according to Ferris, Bongers, & de Goede 2001

protect.sp.pro: plant species categorized into species of concern based on their red list status in Switzerland as well as their protection status in Switzerland and the Canton Zurich (Moser, Gygax, Bäumler, Wyler, & Palese, 2002)

tar.sp: Number of target species (all species recorded in the eleven target plots plus propagules of species applied by hand, resulting in a total of 143 species)

Indi.moist: Plant indicator values for soil moisture according to Landolt et al. 2010

Indi.nutri: Plant indicator values for soil nutrients according to Landolt et al. 2010

Management: level of mowing and manuring frequency (35-40 kg N ha-1 application-1); intensity1 = mowing once and no manure; intensity2 = 2-3 times mowing and no manure; intensity3 = 2-3 times mowing and manuring; intensity4 = 4-5 times mowing and manuring

Tillage: last time of tillage

Land.use.history: for Initial plots = recent agricultural use; for all restored sites = agricultural use before restoration
Mowing	earliest date of biomass harvesting (excluding “Initial”) based on contracts between farmers and the Nature Protection Agency of Zurich

Soil.type: based on soil mapping survey 1991 (before restoration)

Soil.texture: determined by the sediment method (Gee & Bauder 1986) and classified into soil textural classes according to FAO (1990)

pH: soil pH (CaCl2)

Slope: slope gradient classes according to FAO (1990; %)

format: XLS

en: Plant community composition (plant species)

en:
Plot: Plot ID combining treatment code and plot number (in total 55 plots)

Treatment: Three restoration treatments and two reference treatments. Initial=intensively managed grasslands; Harvest only= multiple hay harvest (2-3 times) per years; Topsoil= Topsoil removal (upper 10-30 cm); Topsoil+Propagules= Topsoil removal (upper 10-30 cm) combined with introduction of target plant species; Target= species-rich ancient grasslands

column 3-252: Plant abundance per species (in total 252 plant species) expressed as cover percentage estimation according to van der Maarel (vdM 1979; 7 classes: 1-7); nomenclature according to Lauber & Wagner (1996)

en: Nematode feeding types

en:
Plot: Plot ID combining treatment code and plot number (in total 55 plots)

Treatment: Three restoration treatments and two reference treatments. Initial=intensively managed grasslands; Harvest only= multiple hay harvest (2-3 times) per years; Topsoil= Topsoil removal (upper 10-30 cm); Topsoil+Propagules= Topsoil removal (upper 10-30 cm) combined with introduction of target plant species; Target= species-rich ancient grasslands

He.ecto.a: Herbivorous ectoparasite nematode abundance (individuals per 100g dry soil); classification according to Sieriebriennikov, Ferris, & de Goede, 2014

He.endo.a: Herbivorous endoparasite nematode abundance (individuals per 100g dry soil); classification according to Sieriebriennikov, Ferris, & de Goede, 2014

Ba.a: Bacterivorous nematode abundance (individuals per 100g dry soil); classification according to Sieriebriennikov, Ferris, & de Goede, 2014

Fu.a: Fungivorous nematode abundance (individuals per 100g dry soil); classification according to Sieriebriennikov, Ferris, & de Goede, 2014

Om.a: Omnivorous nematode abundance (individuals per 100g dry soil); classification according to Sieriebriennikov, Ferris, & de Goede, 2014

Ca.a: Carnivorous nematode abundance (individuals per 100g dry soil); classification according to Sieriebriennikov, Ferris, & de Goede, 2014

en: Nematode colonizer-persister classes

en:
Plot: Plot ID combining treatment code and plot number (in total 55 plots)

Treatment: Three restoration treatments and two reference treatments. Initial=intensively managed grasslands; Harvest only= multiple hay harvest (2-3 times) per years; Topsoil= Topsoil removal (upper 10-30 cm); Topsoil+Propagules= Topsoil removal (upper 10-30 cm) combined with introduction of target plant species; Target= species-rich ancient grasslands

CP1.a: C-P (colonizer-persister) class 1 nematode abundance (individuals per 100g dry soil); C-P classification according to Bongers 1990; equivalent to enrichment indicators according to Ferris, Bongers, & de Goede 2001
CP2.a: C-P (colonizer-persister) class 2 nematode abundance (individuals per 100g dry soil); C-P classification according to Bongers 1990; equivalent to stress indicators according to Ferris, Bongers, & de Goede 2001
CP3.a: C-P (colonizer-persister) class 3 nematode abundance (individuals per 100g dry soil); C-P classification according to Bongers 1990
CP4.a: C-P (colonizer-persister) class 4 nematode abundance (individuals per 100g dry soil); C-P classification according to Bongers 1990
CP5.a: C-P (colonizer-persister) class 5 nematode abundance (individuals per 100g dry soil); C-P classification according to Bongers 1990

en: Coordinates of one research plot were wrong. This was corrected in the current version of the file.

en: Nematode families

en:
Plot: Plot ID combining treatment code and plot number (in total 55 plots)

Treatment: Three restoration treatments and two reference treatments. Initial=intensively managed grasslands; Harvest only= multiple hay harvest (2-3 times) per years; Topsoil= Topsoil removal (upper 10-30 cm); Topsoil+Propagules= Topsoil removal (upper 10-30 cm) combined with introduction of target plant species; Target= species-rich ancient grasslands

column 3-41: Nematode abundance (individuals per 100g dry soil) per family; nomenclature according to Bongers 1988

en:
This dataset contains all data on which the following publication below is based.

Paper Citation:
 > Resch, M.C., Schütz, M., Graf, U., Wagenaar, R., van der Putten, W.H., Risch, A.C. 2019. Does topsoil removal in grassland restoration benefit both soil nematode and plant communities? Journal of Applied Ecology 56: 1782-1793.

Please cite this paper together with the citation for the datafile.

 Methods
 Study area and experimental settings
The study was conducted in a nature reserve (Eigental: 47° 27’ to 47° 29’ N, 8° 37’ E, 461 to 507 m a.s.l.) that is located on the Swiss Central plateau close to Zurich airport (Canton Zurich, Switzerland). The mean annual temperature in this area ranges from 8.9 to 10.6 °C, mean annual precipitation from 910 to 1260 mm [10-year average (2007-2017); MeteoSchweiz, 2018]. The main soil types are calcaric to gleyic Cambisol and Gleysols. The reserve was established in 1967 to protect small remnants of oligotrophic semi-natural grasslands (roughly 12 ha). The plant community can be characterized as Molinion and Mesobromion (semi-wet to semi-dry), depending on the site-specific groundwater level and slope inclination (Delarze, Gonseth, Eggenberg, & Vust, 2015). These remnants represent species-rich islands in an otherwise intensively managed agricultural landscape. Semi-natural grasslands covered an area of 60,000 ha in the Canton Zurich in 1939, however, by 2005 only roughly 600 ha remained (Baudirektion Kanton Zürich, 2007). In 1990, the government of Canton Zurich decided to enlarge the nature reserve Eigental. The goal was to incorporate eleven patches of 20 ha adjacent intensively farmed land and transform these patches into semi-natural grasslands. The patches had a different agricultural history, ranging from permanent (no tillage for >50 years) to temporary grassland (as part of crop rotation; last tillage <5 years). On all freshly integrated patches fertilization was stopped in 1992 and from then on biomass was harvested three times a year and removed. After 5 years without noticeable effects on vegetation composition, the Nature Conservation Agency of Canton Zurich decided to increase the restoration efforts. In 1995, a large-scale experiment was initialized to evaluate if certain treatments can facilitate restoration within a reasonable timeframe of 5 to 10 years after treatment implementation.
The three restoration treatments used were:
i. “Harvest only”: Plots are being mowed two to three times a year and the biomass is removed.
ii. “Topsoil”: Topsoil was removed to a depth of 10 to 20 cm, depending on the depth of the O and A horizon, in four randomly selected areas within each of the eleven patches in late autumn 1995. The size of each topsoil removal area depended on individual patch size and was between 2700 and 7000 m2.
iii. “Topsoil+Propagules”: Propagules from target vegetation were added on half of the area where topsoil was removed, using fresh, seed-containing hay originating from a mixture of semi-dry to semi-wet species-rich grasslands of local provenance (within a radius of 7 km). Hay applications were conducted twice in 1995 and 1996. Repeated applications were chosen to account for the low quantity of available plant material per transfer, since area ratio between receptor and donor sites was roughly 1:1. In addition, hand-collected propagules from 15 selected target species of regional provenance (within a radius of 30 km) were equally applied in 1996 and 1997.
“Topsoil” and “Topsoil+Propagules” plots are mowed once a year, and the biomass is removed. Mowing on these plots started five years after the treatment was implemented.
Eleven permanent plots of 5 m x 5 m were randomly established in each treatment to monitor the vegetation development. The experiment was complemented with 11 control plots that represent the initial state of intensively managed grasslands, further referred to as “Initial”, and 11 control plots that represent the targeted state of donor sites for “Topsoil+Propagules”, further referred to as “Target”. Consequently, the experiment consists of 55 plots (5 treatments x 11 replicates). Management of intensively used grasslands includes mowing and fertilizing (manure) between two to five times a year, as well as different tillage regimes (no tillage for >50 years; last time of tillage <5 years).

 Nematode and plant sampling
Soil nematodes were sampled in 2 m x 2 m plots, randomly established at least 2 m away from the vegetation plots. We collected eight soil cores with a 2.2 cm diameter soil core sampler (Giddings Machine Company, Windsor, CO, USA) to a depth of 12 cm (representing the majority of the plant rooting system) in each plot at the beginning of July 2017. The eight cores within each replicate plot were combined, gently homogenized, placed in coolers and transported to the laboratory of NIOO in Wageningen, the Netherlands, within one week. Free-living nematodes were extracted from 200 g of fresh soil using Oostenbrink elutriator (Oostenbrink, 1960) and concentrated, resulting in 6 mL nematode solution. The nematode solution was subdivided into three subsamples, two for morphological identification and quantification, and one for molecular work (not used in this study). For morphological identification and quantification, nematodes were heat-killed at 90 °C and fixed in 4 % formaldehyde solution (final volume 10 mL per subsample). All nematodes in 1 mL of formaldehyde solution were counted, and a minimum of 150 individuals per 1 mL sample (or all if less nematodes were present) were identified to family level using Bongers (1988). We then extrapolated the numbers of each nematode taxa identified to the entire sample and expressed them per 100 g dry soil for further analyses.
We calculated number of nematode taxa and Shannon diversity and assessed nematode community composition. In addition, we classified the nematode taxa into feeding types (herbivores, bacterivores, fungivores, omni-carnivores), structural and functional guilds (Table S4). Structural guilds assign nematode taxa according to life-history traits into five colonizer-persister (C-P) classes, ranging from one (early colonizers of new resources) to five (persisters in undisturbed habitats; Bongers 1990). C-P classes can be categorized as indicators for nutrient-enriched (C-P1), stressed (C-P2) and structured (C-P3 + C-P4 + C-P5) soil conditions (Ferris, Bongers, & de Goede, 2001). Functional guilds assign nematode taxa according to their C-P classification combined with their feeding habits (Ferris, Bongers, & de Goede, 2001). Based on the structural and functional guild classification we calculated five additional indices to assess soil nutrient status, disturbance and food web characteristics using NINJA (Sieriebriennikov, Ferris, & de Goede, 2014). 1) The Maturity index indicates the degree of different environmental perturbations (e.g., tillage, nutrient enrichment, pollution) and is used to monitor colonization and subsequent succession after disturbances (Bongers, 1990). 2) The ratio between the Plant Parasite (C-P of herbivorous nematodes only) to Maturity index is used to monitor the recovery of disturbed habitats incorporating information of life-history traits for all feeding types (Bongers, van der Meulen, & Korthals, 1997). 3) The Enrichment index indicates nutrient-enriched soils and agricultural management practices (Ferris, Bongers, & de Goede, 2001). 4) The Structure index provides information about the succession stage of the soil food web and therefore correlates with the degree of maturity of an ecosystem (Ferris, Bongers, & de Goede, 2001). 5) The Channel index provides information about the predominant decomposition pathways, where higher values stand for a higher proportion of energy transformed through the slow fungal decomposition channel (Ferris, Bongers, & de Goede, 2001). In addition, the Structure and Enrichment indices can be displayed in a biplot where nematode assemblages are plotted along a structure (x-axis) and enrichment (y-axis) trajectory (increasing index values). Each biplot quadrat reflects different levels of disturbance, soil nutrient pools and decomposition pathways (Ferris, Bongers, & de Goede, 2001).
The plant surveys were conducted on the 25 m2 permanent plots in June 2017. Plant species cover was visually assessed according to the semi-quantitative cover-abundance scale of Braun-Blanquet (1964; nomenclature: Lauber & Wagner, 1996). We calculated number of species and Shannon diversity, and assessed plant community composition. We also counted the number of target species (all species recorded in the eleven target plots plus propagules of species applied by hand, resulting in a total of 143 species) and categorized plant species into species of concern based on their red list status in Switzerland as well as their protection status in Switzerland and the Canton Zurich (Moser, Gygax, Bäumler, Wyler, & Palese, 2002). Furthermore, we calculated indicator values for soil moisture and soil nutrients for each species according to Landolt et al. (2010).

 References
Baudirektion Kanton Zürich (2007). 10 Jahre Naturschutz-Gesamtkonzept für den Kanton Zürich 1995-2005 – Stand der Umsetzung. Zürich: Baudirektion Kanton Zürich.
Bongers, T. (1988). De nematoden van Nederland. Utrecht: Stichting Uitgeverij Koninklijke Nederlandse Natuurhistorische Vereniging.
Bongers, T. (1990). The maturity index: an ecological measure of environmental disturbance based on nematode species composition. Oecologia, 83, 14-19. doi:10.1007/BF00324627
Bongers, T., van der Meulen, H., & Korthals, G. (1997). Inverse relationship between the nematode maturity index and plant parasite index under enriched nutrient conditions. Applied Soil Ecology, 6, 195-199. doi:10.1016/S0929-1393(96)00136-9
Braun-Blanquet, J. (1964). Pflanzensoziologie, Grundzüge der Vegetationskunde (3rd ed.). Wien: Springer.
Delarze, R., Gonseth, Y., Eggenberg, S., & Vust, M. (2015). Lebensräume der Schweiz: Ökologie - Gefährdung - Kennarten (3rd ed.). Bern: Ott.
Ferris, H., Bongers, T., & de Goede, R.G.M. (2001). A framework for soil food web diagnostics: extension of the nematode faunal analysis concept. Applied Soil Ecology, 18, 13-29. doi:10.1016/S0929-1393(01)00152-4
Landolt, E., Bäumler, B., Erhardt, A., Hegg, O., Klötzli, F., Lämmler, W., … Wohlgemuth, T. (2010). Flora indicativa. Ecological indicator values and biological attributes of the Flora of Switzerland and the Alps (2nd ed.). Bern: Haupt.
Lauber, K., & Wagner, G. (1996). Flora Helvetica. Flora der Schweiz. Bern: Haupt.
MeteoSchweiz (2018). Klimabulletin Jahr 2017, Zürich: MeteoSchweiz.
Moser, D., Gygax, A., Bäumler, B., Wyler, N., & Palese, R. (2002) Rote Liste der gefährteten Farn- und Blütenpflanzen der Schweiz. Bern: BUWAL.
Oostenbrink, M. (1960). Estimating nematode populations by some selected methods. In N.J. Sasser & W.R. Jenkins (Eds.), Nematology (pp. 85-101). Chapel Hill: University of North Carolina Press.
Sieriebriennikov, B., Ferris, H., & de Goede, R.G.M (2014). NINJA: An automated calculation system for nematode-based biological monitoring. European Journal of Soil Biology, 61, 90-93. doi:10.1016/j.ejsobi.2014.02.004

diversity

display_name: diversity

grassland

display_name: grassland

long-term-experiment

display_name: long-term-experiment

nematodes

display_name: nematodes

restoration

display_name: restoration

topsoil-removal

display_name: topsoil-removal

vegetation

display_name: vegetation

educ

en: Education, culture and sport

en: Grassland restoration: nematodes and plant communities


modeling-snow-failure-with-dem

title_for_slug: modeling-snow-failure-with-dem

en: snow-failure snow-avalanche discrete-element-modelling snow-micro-mechanics snow

en: Python file

en: Python file to reproduce the figures in Bobillier et al. (2019)

mimetype: text/x-python

en: Read-me

media_type: application/pdf

en: The read-me file contains a description of all the data provided in the zip file.

format: PDF

mimetype: application/pdf

en: Dataset

media_type: application/zip

en: The zip file contains the data underlying the figures presented in Bobillier et al. (2019). The figures can be reproduced with the python file, which is also provided.

format: ZIP

mimetype: application/zip

en: This data set includes the modeling results described in the research article by Bobiller et al. (2020). All the figures in the article can be reproduced with the data provided.

discrete-element-modelling

display_name: discrete-element-modelling

snow-avalanche

display_name: snow-avalanche

snow-failure

display_name: snow-failure

snow-micro-mechanics

display_name: snow-micro-mechanics

envidat

en: EnviDat: the Environmental Data Portal

en: EnviDat is the WSL data portal providing unified and managed access to environmental monitoring and research data. The portal has the capability to host and publish data sets. Through the sharing of unique and high-value data sets worldwide, EnviDat fosters research and contributes to advancing environmental science.

en: Modeling snow failure with DEM


snow-drift-station-flowcapt

maintainer: Benjamin Walter

title_for_slug: snow-drift-station-flowcapt

en: snow-blowing snowdrift flowcapt raclets acoustic-mass-flux wind-speed

en: minimum mass flux

en: The data file contains the minimum mass flux of 10 minute intervalls. The duration of one cycle was 1 min and contained 6s observations and 54 s stand-by. The time is given in UTC+1.

en: average wind speed

en: The data file contains the average wind speed in 10 minute intervalls. The duration of one cycle was 1 min and contained 6s observations and 54 s stand-by. The time is given in UTC+1.

en: maximum mass flux

en: The data file contains the maximum mass flux of 10 minute intervalls. The duration of one cycle was 1 min and contained 6s observations and 54 s stand-by. The time is given in UTC+1.

en: maximum wind speed

en: The data file contains the maximum wind speed in 10 minute intervalls. The duration of one cycle was 1 min and contained 6 s observations and 54 s stand-by. The time is given in UTC+1.

en: avergaged mass flux

en: Average mass flux of 10 min measurement interval. The Cycle duration is 1 min which includes 6 s observation and 54 s stand-by. The time is given in UTC+1.

en: minimum wind spees

en: The data file contains the minimum wind speed in 10 minute intervalls. The duration of one cycle was 1 min and contained 6s observations and 54 s stand-by. The time is given in UTC+1.

en: The FlowCapt is an ultra-robust instrument measuring solid particle acoustic mass - flux intensities (g m‾ ² s‾ ¹) and wind speeds (m s‾ ¹). It was deployed at Gotschnagrat (LON: 46.859 LAT: 9.849). The vertical tube with a length of 1 m monitors snowdrift and snow-blowing; and is mounted at a height between 0.1 an 1.1 m above snow surface. The time within each data set is given in UTC+1.

acoustic-mass-flux

display_name: acoustic-mass-flux

flowcapt

display_name: flowcapt

raclets

display_name: raclets

snow-blowing

display_name: snow-blowing

snowdrift

display_name: snowdrift

wind-speed

display_name: wind-speed

en: Snow Drift Station - Flowcapt


disdrometer-data-gotschnagrat

title_for_slug: disdrometer-data-gotschnagrat

en: raclets precipitation visibility disdrometer hydrometeors

en: Disdrometer Data Gotschnagrat

en: The zip folder contains one netCDF file per day for Gotschnagrat from February 11 to March 27 2019 with a temporal resolution of 30 seconds (time in UTC). The naming convention for the files is Gotschna_YYYYMMDD.nc and the files include the following: characteristics of hydrometeors (e.g. diameter, fall velocity, amount per diameter class,...), precipitation rate, radar reflectivity, visibility range, weather codes and instrument performance.

en: A laser optical disdrometer (Parsivel² , OTT Hydromet) was deployed at Gotschnagrat (LON: 9.849, LAT: 46.859) to measure hydrometeors by extinction when passing a laser beam. The instrument can classify eight different kinds of precipitation, including rain, hail, snow, drizzle, and hybrid forms. The dataset contains information on precipitation amount and type for the period of February 11 to March 27 2019 at Gotschnagrat.

disdrometer

display_name: disdrometer

hydrometeors

display_name: hydrometeors

precipitation

display_name: precipitation

visibility

display_name: visibility

educ

en: Education, culture and sport

envidat

en: EnviDat: the Environmental Data Portal

en: EnviDat is the WSL data portal providing unified and managed access to environmental monitoring and research data. The portal has the capability to host and publish data sets. Through the sharing of unique and high-value data sets worldwide, EnviDat fosters research and contributes to advancing environmental science.


wind-crust-formation-snowmicropen-data

title_for_slug: wind-crust-formation-snowmicropen-data

en: snowmicropen snow smp wind-packing wind-crust

en: Processing scripts

en: Contains the R-scripts used to process the SMPs

en: README

media_type: text/plain

en: Describes the dataset in detail

format: TXT

mimetype: text/plain

en: SMP data

en: Links to the dataset

en:
This dataset contains the SnowMicroPen (SMP) data from 38 wind tunnel experiments on wind-packing / wind crust formation. These experiments were performed in the winters 2015/16 and 2016/17 and include more than 1000 SMP measurements.

The SMPs are organized per experiment. Each experiment subfolder contains the processed SMP profiles and some additional files.

Please refer to the README for more details on the data. The processing scripts are available for download as well. The scripts are mainly provided as documentation and would need to be adjusted to be used.

This dataset is the basis of the following publication:

Sommer C.G., Lehning M., & Fierz C. (2017). Wind tunnel experiments: Saltation is necessary for wind-packing. Journal of Glaciology, 63(242), 950-958. doi:10.1017/jog.2017.53

smp

display_name: smp

snow

display_name: snow

snowmicropen

display_name: snowmicropen

wind-crust

display_name: wind-crust

wind-packing

display_name: wind-packing

en: Wind crust formation: SnowMicroPen data


maintainer: Rebecca Mott

en: turbulent-sensible-heat-fluxes snow-patch patchy-snow-covers eddy-covariance-measurements dischmex

en: Eddy-covariance measurement with an ultrasonic anemometer (DA-600, Kaijo-Denki) in 0.3 m on a daily basis.

en: Eddy-covariance measurement with an ultrasonic anemometer (CSAT3, Campbell Scientific, Inc.) in 2.2 m.

en: Eddy-covariance measurement with an ultrasonic anemometer (CSAT3, Campbell Scientific, Inc.) in 2.6 m.

en: Eddy-covariance measurement with an ultrasonic anemometer (CSAT3, Campbell Scientific Inc.) in 3.2 m on a daily basis.

en: Eddy-covariance measurement with an ultrasonic anemometer (R.M. Young Company) in 3.3 m.

en: Eddy-covariance measurement with an ultrasonic anemometer (R.M. Young Company) in 3.5 m.

en: Output explanation

en: Eddy-covariance measurement with an ultrasonic anemometer (R.M. Young Company) in 0.7 m.

en: Gletschboden setup

en: Background map based on data © 2014/2016 swisstopo (JD100042)

mimetype: image/png

en: Eddy-covariance measurement with an ultrasonic anemometer (CSAT3, Campbell Scientific Inc.) in 1.7 m on a daily basis.

en:
This dataset contains eddy-covariance measurements in the ablation period of 2014-2016. Measurements were taken from two turbulence towers over a long-lasting snow patch, which are 5 m apart from each other (2014 and 2015). The turbulence towers were equipped with two YOUNG ultrasonic anemometers mounted 0.7 m (in 2014) and 3.3 m (in 2015) above snow-free ground, two ultrasonic anemometers (CSAT3, Campbell Scientific, Inc.) mounted at 2.6 m (in 2014) and 2.2 m (in 2015) above snow-free ground and one anemometer (DA-600, Kaijo Denki) mounted at 0.3 m above snow surface.

The measurement setup changed in 2016 and includes a measurement above the snow-free ground in upwind direction (Swiss coordinates: 790191/176689). The measurement tower is equipped with one ultrasonic anemometer (CSAT3, Campbell Scientific, Inc.) in 3.3 m above the snow-free ground. Additionally, one measurement tower is installed above the long-Lasting snow patch and equipped with the same setup as 2015.

Turbulence data were sampled at a frequency of 20 Hz. The processing of the data to quality controlled fluxes has been done with the Biomicrometeorology flux software (Thomas et al., 2009). The program applies plausibility tests and a despiking test after Vickers and Mahrt (1997) on the measured data. The routine further applies a time-lag correction and considers the deployment (e.g. the sonic azimuth). A frequency response correction (Moore, 1986) is done and a three-dimensional rotation is performed. Finally, quality assurance/quality control (QA/QC) flags after Foken et al., (2004) are issued and fast Fourier transform power and co-spectra are calculated. The change in snow height is considered in the post-processing for every measurement day. The turbulence data were averaged to 30 minute intervals.

dischmex

display_name: dischmex

eddy-covariance-measurements

display_name: eddy-covariance-measurements

patchy-snow-covers

display_name: patchy-snow-covers

snow-patch

display_name: snow-patch

turbulent-sensible-heat-fluxes

display_name: turbulent-sensible-heat-fluxes

en: DISCHMEX - Impact of extreme land-surface heterogeneity on micrometeorology over spring snow-cover


maintainer: Thorsten Bartels-Rausch

en: interface chemistry snow arctic ozone antarctica laboratory-study alpine metamorphism reaction brine solution heterogeneous

en: Matlab Script to view and plot data

en: Data Figure 1

en: Data of published Figure

en: Readme file

en: This file shows data, code, and plots.

en: Matlab Data

en: All data as Matlab mat file

en: data list

en: Lists file names, snow sample properties, and flow settings for each data-set.

en: Acetone data

en:
intensity of the acetone-water-cluster and water-cluster (as derived by a chemical ionisation mass spectrometer) with time in the carrier gas downstream of packed bed flow tube holding the snow sample or the by-pass, respectively. Time "0" denotes the time when the mass spectrometer data acquisition was started. When the carrier
gas with acetone was passed via the packed bed flow tube, that is switched from by-pass to pass the snow, acetone concentration in the carrier gas rapidly dropped due to surface adsorption and slowly recovered as the adsorption sites saturate. From this, the surface area can be obtained.

en: Ozone data

en: Trend of ozone mixing ratio ( _O3mixingRatio_ppb_ with time ( _time_s_ ) as measured downstream of the snow sample during the experiments. Time "0" denotes the time when data acquisition with ozone analyser was started. And, ozone (_nOzone_molec_per_s_ ) loss for each experiment with duration of ozone exposure ( _duration_s_ ) of ozone exposure, as derived from the ozone trends shown above. Time "0" denotes the time when the carrier gas with ozone was passed via the snow sample.

en: Earth’s snow cover is very dynamic on diurnal time scales. The changes to the snow structure during this metamorphism have wide ranging impacts such as on avalanche formation and on the capacity of surface snow to exchange trace gases with the atmosphere. Here, we investigate the influence of dry metamorphism, which involves fluxes of water vapor, on the chemical reactivity of bromide in the snow. For this, the heterogeneous reactive loss of ozone at a concentration of 5-6E12 molecules cm-3 is investigated in artificial, shock-frozen snow samples doped with 6.2 uM sodium bromide and with varying metamorphism history. The oxidation of bromide in snow is one reaction initiating polar bromine releases and ozone depletions.

alpine

display_name: alpine

antarctica

display_name: antarctica

arctic

display_name: arctic

brine

display_name: brine

chemistry

display_name: chemistry

heterogeneous

display_name: heterogeneous

interface

display_name: interface

laboratory-study

display_name: laboratory-study

metamorphism

display_name: metamorphism

ozone

display_name: ozone

reaction

display_name: reaction

solution

display_name: solution

educ

en: Education, culture and sport

envidat

en: EnviDat: the Environmental Data Portal

en: EnviDat is the WSL data portal providing unified and managed access to environmental monitoring and research data. The portal has the capability to host and publish data sets. Through the sharing of unique and high-value data sets worldwide, EnviDat fosters research and contributes to advancing environmental science.

en: Data set on bromide oxidation by ozone in snow during metamorphism from laboratory study


en: sdms soil precipitation ground-temperature aridity frost-change-velocity

media_type: image/tiff

en: Mean diurnal range modified from CHELSA

format: TIFF

en: Precipitation seasonality from CHELSA

en: Soil pH modified from SoilGrids

en: Soil organic carbon content modified from SoilGrids

en: Frost change frequency modified from CHELSA

en: Soil clay content modified from SoilGrids

en: Precipitation of driest quarter modified from CHELSA

en: Mean annual temperature modified from CHELSA

en: The dataset contains seven environmental layers (average annual temperature, aridity [annual precipitation divided by annual potential evapotranspiration], frost change frequency, precipitation in the driest quarter, mean diurnal temperature range, and precipitation seasonality) modified from CHELSA (https://chelsa-climate.org/) and three soil layers (soil organic matter content, pH water, and clay content) modified from SoilGrids (https://soilgrids.org/).

aridity

display_name: aridity

frost-change-velocity

display_name: frost-change-velocity

ground-temperature

display_name: ground-temperature

precipitation

display_name: precipitation

sdms

display_name: sdms

soil

display_name: soil

en: Environmental layers for SDM simulations (GDPlants)


rorcc

maintainer: Tobias Wechsler

title_for_slug: rorcc

en: hydropower design-discharge climate-change environmental-flow run-of-river-power-plants

en: RoRCC

en:
The dataset “RoRCC” consists of simulation-based results on climate change impacts on Alpine RoR power production; it is based on 21 Swiss RoR power plants, with a total production of 5.9 TWh a-1. The dataset contains the following information:
-	metadata on the RoR power plants under consideration
-	annual and seasonal production scenarios differentiated into three emission scenarios (RCP2.6, RCP4.5, RCP8.5) and three future periods (T1: 2020–2049, T2: 2045–2074, T3: 2070–2099)
-	annual and seasonal production loss due to environmental flow requirements
-	annual and seasonal the technical increase potential (via design discharge optimisation)
-	annual and seasonal changes in the hydrological cycle for the three emission scenarios (RCP2.6, RCP4.5, RCP8.5) and three future periods (T1: 2020–2049, T2: 2045–2074, T3: 2070–2099)

en:
The dataset "RoRCC" consists of simulation-based results on climate change impacts on Alpine RoR power production; it is based on 21 Swiss RoR power plants, with a total production of 5.9 TWh a-1. The dataset contains the following information:
1) metadata on the RoR power plants under consideration, 2) annual and seasonal production potential scenarios under into three emission scenarios (RCP2.6, RCP4.5, RCP8.5) and three future periods (T1: 2020–2049, T2: 2045–2074, T3: 2070–2099), 3) annual and seasonal streamflow scenarios, 4) annual and seasonal production loss due to environmental flow requirements, 5) annual and seasonal the technical increase potential (via design discharge optimisation) and 6) annual and seasonal changes in the hydrological cycle.

climate-change

display_name: climate-change

design-discharge

display_name: design-discharge

environmental-flow

display_name: environmental-flow

hydropower

display_name: hydropower

run-of-river-power-plants

display_name: run-of-river-power-plants


en: precipitation-variability orographic-precipitation cosmo-wrf complex-terrain snow-precipitation

en: Snow depth maps over Dischma valley for 26 January 2016 and 9 March 2016 and snow depth differences between the two dates.

en: StationData

en:
The data presented here corresponds to the publication "Spatial variability in snow precipitation and accumulation in COSMO-WRF simulations and radar estimations over complex terrain" (Gerber et al., 2018a), which investigates the precipitation variability of snow precipitation in the central northern part of the Grisons (CH) and the publication "The importance of near-surface winter precipitation processes in complex alpine terrain" (Gerber et al., 2018b). The dataset contains:

* WRFsimulations: WRF simulation output for simulations with 4x (14x) terrain smoothing with an output timestep of 30 min/5 min and horizontal grid spacings of 1350 m, 450 m, 150 m and 50 m (currently: data available upon request).
* StationData: Meteorological station data of 18 meteorological stations in the central northern part of the Grisons with 30 minute resolution for the period 1 January 2016 till 1 May 2016.
* ADS80data: Photogrammetrically determined snow depth distribution data over the Dischma valley for the 26 January 2016 and 9 March 2016. Snow heights are corrected for buildings, vegetation (> 1m), outliers, and pixles, which are obivously snow-free on the pictures (Bühler et al., 2015). In addition the snow depth differences (snow depth on 9 March 2016 minus snow depth on 26 January 2016) are provided.

For more details about the simulation and observation data, see Gerber et al., 2018 and Gerber and Sharma (2018).

Publications:

Bühler, Y., Marty, M., Egli, L., Veitinger, J., Jonas, T., Thee, P., and Ginzler, C.: Snow depth mapping in high-alpine catchments using digital photogrammetry. Cryosphere, 9, 229–243, doi:10.5194/tc-9-229-2015, 2015.

Gerber, F., Besic, N., Sharma, V., Mott, R., Daniels, M., Gabella, M., Berne, A., Germann, U., and Lehning, M.: Spatial variability in snow precipitation and accumulation in COSMO-WRF simulations and radar estimations over complex terrain, The Cryosphere, 12, 3137–3160, doi:10.5194/tc-12-3137-2018, 2018.

Gerber, F., Mott, R. and Lehning, M.: The importance of near-surface winter precipitation processes in complex alpine terrain, Journal of Hydrometeorology, accepted, 2018.

Gerber, F., and Sharma, V.: Running COSMO-WRF on very-high resolution over complex terrain. Laboratory of Cryospheric Sciences CRYOS, École Polytechnique Fédérale de Lausanne EPFL, Lausanne, Switzerland. doi:10.16904/envidat.35, 2018.

complex-terrain

display_name: complex-terrain

cosmo-wrf

display_name: cosmo-wrf

orographic-precipitation

display_name: orographic-precipitation

precipitation-variability

display_name: precipitation-variability

snow-precipitation

display_name: snow-precipitation

educ

en: Education, culture and sport

envidat

en: EnviDat: the Environmental Data Portal

en: EnviDat is the WSL data portal providing unified and managed access to environmental monitoring and research data. The portal has the capability to host and publish data sets. Through the sharing of unique and high-value data sets worldwide, EnviDat fosters research and contributes to advancing environmental science.

en: DISCHMEX - High-resolution WRF simulations in complex alpine terrain and station measurements


en: pest-invasion southern-switzerland castanea-sativa torymus-sinensis dryocosmus-kuriphilus population-evolution biological-control

en:
Data has been collected from summer 2011 to 2016.

Main reasons for NA values:

-	D. kuriphilus still not arrived in the site.
-	No galls were found during the field sampling.
-	Site has not been sampled in the correspondent year because of time, weather or money constrains.
-    T. sinensis not found in the galls

Variable description:

Year = sampling year.

Coordinate, elevation, slope, aspect = average site characteristics.

ts_arrival = T. sinensis arrival year.

dk_arrival = D. kuriphilus arrival year.

dk_parasitism_by_ts = D. kuriphilus parasitism rate by T. sinensis. Formula: (no. of T. sinensis/ no. of D. kuriphilus chambers) * 100 at gall level.

dk_infestation = Percentage of attacked buds. Formula: no. of attacked buds / no. of total developed buds*100 (as presented in Kotobuki et al. 1985) at shoot level.

ts_per_bud = number of alive T. sinensis individual calculated at bud level.

dk_per_bud = number of alive D. kuriphilus individual calculated at bud level.

dk_pot_per_bud = overall number (dead and alive) D. kuriphilus individual calculated at bud level.

total_gall_nr_per_bud = average gall number per bud.

chamber_per_galls = average D. kuriphilus chamber number per gall.

For details see the submitted publication with the same name

References
Kotobuki, K., Mori, K. and Sato, Y. (1985) ‘2 methods to estimate the tree damage by chestnut gall wasp Dryocosmus-kuriphilus’, Bulletin of the fruit tree research station A (Yatabe), 2(12), pp. 29–36.

en:
This dataset contains the population evolution of a pest and its biocontrol agent in terms of presence proportion at gall level and absolute number of insects.

The study area extends from the Cuneo region (Piedmont, Italy) to southern Switzerland. In order to provide a complete range of data covering the entire process from the pest arrival to complete biological control by its natural enemy T. sinensis, a space-for-time substitution approach has been adopted so as to create a temporal gradient of the epidemic stages over the whole study area.

The southernmost Swiss sites roughly represent the arrival and establishment of the pest without the presence of the natural enemy, the central ones the early epidemic stage and the epidemic peak, whereas the northern ones the end of the epidemic with the beginning of the biocontrol. The Italian ones represent the beginning of the equilibrium between the two population as well as the situation with stable T. sinensis populations on the long term.

These data are used in the paper entitled: Torymus sinensis local and regional early population dynamics in the Insubrian and Piedmont regions

biological-control

display_name: biological-control

castanea-sativa

display_name: castanea-sativa

dryocosmus-kuriphilus

display_name: dryocosmus-kuriphilus

pest-invasion

display_name: pest-invasion

population-evolution

display_name: population-evolution

southern-switzerland

display_name: southern-switzerland

torymus-sinensis

display_name: torymus-sinensis


aerosol-data-weissfluhjoch

title_for_slug: aerosol-data-weissfluhjoch

en: ice-nucleation raclets aerosol linda inp particles

en: OPC-N3 Dataset Weissfluhjoch

en: Particle concentrations in a size range of 0.35 μm to 40 μm were measured with an OPC-N3 (Alphasense, Inc.) in a resolution of 20 seconds (time in UTC). PM₁, PM₂,₅ and PM₁₀ were directly derived.

en: APS Dataset Weissfluhjoch

en: The APS file contains aerosol data (total number concentration, median, mean, geometric mean, mode and geometric standard deviation of the size distribution) for aerodynamic diameters between 0.5 μm and ~20 μm as well as instrument characteristics, in a temporal resolution of 20 seconds, with time in UTC, measured at Weissfluhjoch.

mimetype: application/x-netcdf

en: SMPS Dataset Weissfluhjoch

en: The SMPS file contains aerosol data (particle concentration, total number concentration, median, mean, geometric mean, mode and geometric standard deviation of the size distribution) in a size range between 12 and 460 nm (electrical mobility diameter) as well as instrument characteristics (sample temperature, pressure, relative humidity, mean free path and gas viscosity) in a temporal resolution of 2 minutes, with time in UTC, measured at Weissfluhjoch.

en: LINDA Dataset Weissfluhjoch

en: The LINDA file contains information on temperature (in °C) dependent frozen fractions, number of Ice Nuleating Particles (INP, per standard liter) and differential INP concentrations (per standard liter and Kelvin).

en: OPC-GT526S Dataset Weissfluhjoch

en: Particle counts were recorded in six size channels (0.3, 0.5, 0.8, 1.5, 2.5 and 5 μm) in a resolution of 20 seconds (time in UTC) by a particle counter (GT-526S, Met One Instruments, Inc.).

en:
Aerosol properties were measured between February 8 and March 31 2019 at the measurement site Weissfluhjoch (LON: 9.806475, LAT: 46.832964). Optical and aerodynamic particle counters, as well as a scanning mobility particle size spectrometer and an ice nuclei counter were deployed to report particle concentrations and size distributions in fine (10-1000 nm) and coarse mode (> 1000 nm), cloud condensation nuclei concentrations (CCNCs), and ice nuclei particle concentrations (ICNCs). The ambient particles were transported via a heated inlet to be distributed to the particle detecting devices inside the setup room.

Optical Particle Counter (OPC): Light scattering of a diode laser beam caused by travelling particles is used in the both, the OPC-N3 (0.41 - 38.5 μm) and GT-526S (0.3 – 5 μm), to determine their size and number concentration. For the OPC-N3, particle size spectra and concentration data are used afterwards to calculate PM₁, PM₂,₅ and PM₁₀ (assumptions: particle density: 1.65 g cmˉ³, refractive index: 1.5+i0).

Aerodynamic Particle Sizer (APS): The APS (3321, TSI Inc.) measured the particle size distribution for aerodynamic diameters between 0.5 μm and ~20 μm by the particle’s time-of-flight and light-scattering intensity (assumptions: particle density 1 g cmˉ³).

Scanning Mobility Particle Size Spectrometer (SMPS): Particle number concentrations in a size range between 12 and 460 nm (electrical mobility diameter) were measured at Davos Wolfgang, using a Scanning Mobility Particle Sizer Spectrometer (SMPS 3938, TSI Inc.). The classifier (3082, TSI Inc.) was equipped with a neutralizer (3088, TSI Inc.) and a differential mobility analyzer working with negative polarity (3081, TSI Inc.). The size selected particles were counted by a water-based condensation particle counter (3787 TSI Inc.). The TSI AIM software was used to provide particle size distributions by applying multiple charge and diffusion loss corrections (assumptions: particle density 1 g cmˉ³).

Coriolis μ and LINDA: A microbial air sampler (Coriolis μ, bertin Instruments) was used to collect airborne particles for investigating their ice nucleating ability with a droplet freezing device. Particles larger than 0.5 μm were drawn with an air flow rate of up to 300 l min‾¹ into the cone and centrifuged into the wall of the cone due to the forming vortex. The liquid sample was transferred into the LED based Ice Nucleation Detection Apparatus (LINDA, University of Basel) to study heterogeneous ice formation (immersion freezing mode) of ambient airborne particles.

aerosol

display_name: aerosol

ice-nucleation

display_name: ice-nucleation

inp

display_name: inp

linda

display_name: linda

particles

display_name: particles

raclets

display_name: raclets

en: Aerosol Data Weissfluhjoch


en: mass-flux antarctica drifting-snow terrestrial-laser-scanning snowmicropen meteorological-data

en: README

en: Describes the dataset in detail

en: Dataset

en: Links to the dataset

en:
This dataset contains the data acquired during the expedition to Princess Elisabeth Antarctica Station in December 2016 and January 2017. The dataset consits of meterorological data, drifting snow mass flux data, SnowMicroPen data and Terrestrial Laser Scanning data.

Please refer to the README for more information about the data.

This dataset is the basis of the following publication:

Sommer, C. G., Wever, N., Fierz, C., and Lehning, M.: Wind-packing of snow in Antarctica, The Cryosphere Discuss., https://doi.org/10.5194/tc-2018-36, in review, 2018.

antarctica

display_name: antarctica

drifting-snow

display_name: drifting-snow

mass-flux

display_name: mass-flux

meteorological-data

display_name: meteorological-data

snowmicropen

display_name: snowmicropen

terrestrial-laser-scanning

display_name: terrestrial-laser-scanning

educ

en: Education, culture and sport

envidat

en: EnviDat: the Environmental Data Portal

en: EnviDat is the WSL data portal providing unified and managed access to environmental monitoring and research data. The portal has the capability to host and publish data sets. Through the sharing of unique and high-value data sets worldwide, EnviDat fosters research and contributes to advancing environmental science.

en: Expedition to Princess Elisabeth Antarctica Station, 2016/2017