maintainer: Anita Risch

en: vertebrates ecosystem-multifunctionality swiss-national-park ecosystem-functions ecosystem-coupling subalpine grassland grazing exclusion invertebrates

en: Datafile

en:
ID: Plot ID composited of Fence & Treatment number (90 plots total)

Grassland: Name of the six subalpine grasslands where the exclosure setups were erected. In large grasslands (GRIM, MING, STAB), we established four hierarchical exclosure setups (two in short- and two in tall-grass vegetation), whereas in the small grasslands (BOTS, DADA, MARG) only one setup was erected per vegetation type. BOTS = Val dal Botsch, GRIM = Grimmels, MING = Val Mingér, STAB = Alp Stabelchod, DADA = Stabelchod da daint, MARG = Margunet

Fence: Number of exclosure setup (1 - 18). Note that uneven numbers (1,3,5,7,9,11,13,15,17) represent setups established on short-grass vegetation, even numbers (2,4,6,8,10,12,14,16,18) setups established in tall-grass vegetation
Vegetation	Vegetation type, 1 = short-grass vegetation, 2 = tall-grass vegetation; (9 replicate fences on each vegetation type)
Treatment	Progressive animal exclusion, numbers indicate which animals had access to the plot: 1 = Large/medium/small mammals, and invertebrates have access ("L/M/S/I"), 2 = Medium/small mammals, and invertebrates have access (“M/S/I”), 3 = Small mammals and invertebrates have access (“S/I”), 4 = Invertebrates have access (“I”), 5 = No animals have access (“None”)

Bulkden:	Fine fraction soil bulk density (g cm-3)

Soil.moist	: Soil moisture (%)

Soil.org.C: Rhizosphere soil C content (%)

pH: Soil pH

Nema.PC1: First principal component (PC1) of principal component analysis (PCA) of the soil nematode community composition

Nema.PC2: Second principal component (PC2) of principal component analysis (PCA) of the soil nematode community composition

Micro.PC1: 	First principal component (PC1) of principal component analysis (PCA) of the soil microbial community composition

Micro.PC2: Second principal component (PC2) of principal component analysis (PCA) of the soil microbial community composition

Abv.Inv.PC1: First principal component (PC1) of principal component analysis (PCA) of the aboveground-dwelling invertebrate community composition

Abv.Inv.PC2: Second principal component (PC2) of principal component analysis (PCA) of the aboveground-dwelling invertebrate community composition

Arthr.PC1: First principal component (PC1) of principal component analysis (PCA) of the soil arthropod community composition

Arthr.PC2: Second principal component (PC2) of principal component analysis (PCA) of the soil arthropod community composition

Plant.PC1: First principal component (PC1) of principal component analysis (PCA) of the plant community composition

Plant.PC2: Second principal component (PC2) of principal component analysis (PCA) of the plant community composition

Root.biomass: Root biomass (g m-2)

Micro.biomass	: Microbial biomass carbon (mg kg-1)

Plant.richness: Plant species richness

Plant.N: Plant tissue nitrogen content (%)

Soil.N.min: Soil net nitrogen mineralisation (kg ha-1 season)

Soil.resp: Soil carbon dioxide emissions (soil respiration; g m-2 h-1)

format: XLS

en:
This dataset contains all data on which the following publication below is based.

Paper Citation:
 > Risch AC, Ochoa-Hueso R, van der Putten WH, Bump JK, Busse MD, Frey B, Gwiazdowicz DJ, Page-Dumroese DS, Vandegehuchte ML, Zimmermann S, Schütz M. Size-dependent loss of aboveground animals differentially affects grassland ecosystem coupling and functions. 2018. Nature Communications 9: 3684. [doi: 10.1038/s41467-018-06105-4](https://doi.org/10.1038/s41467-018-06105-4).

Please cite this paper together with the citation for the datafile.

#Methods
Study sites
The experimental exclosure setups were installed within the SNP (IUCN category Ia preserve; Dudley 2008), in south-eastern Switzerland. The park covers 172 km2 of forests and subalpine and alpine grasslands along with scattered rock outcrops and scree slopes. The entire area has been protected from human impact (no hunting, fishing, camping or off-trail hiking) since 1914. Large, fairly homogenous patches of short- and tall-grass vegetation, which originate from different historical management and grazing regimes, cover the park’s subalpine grasslands entirely. Short-grass vegetation developed in areas where cattle used to rest (nutrient input) prior to the park’s foundation (14th century to 1914) (Schütz and others 2003, 2006) and is dominated by lawn grass species such as Festuca rubra L., Briza media L. and Agrostis capillaris L. (Schütz and others 2003, 2006). Today, this vegetation type is intensively grazed by diverse vertebrate and invertebrate communities that inhabit the park and consume up to 60% of the available biomass (Risch and others 2013). Tall-grass vegetation developed where cattle formerly grazed, but did not rest, and is dominated by rather nutrient-poor tussocks of Carex sempervirens Vill. and Nardus stricta L. (Schütz and others 2003, 2006). This vegetation type receives considerably less grazing, with only roughly 20% of the biomass consumed (Risch and others 2013). Consequently, the two vegetation types together represent a long-term trajectory of changes in grazing regimes. Underlying bedrock of all grasslands is dolomite, which renders these grasslands rather poor in nutrients regardless of former and current land-use regimes.
Experimental design
To progressively exclude aboveground vertebrate and invertebrate animals, we established 18 size-selective exclosure setups (nine in short-grass, nine in tall-grass vegetation) distributed over six subalpine grasslands across the SNP (Risch and others 2013, 2015). Elevation differences of exclosure locations did not exceed 350 m (between 1975 and 2300 m a.s.l.). The exclosures were established immediately after snowmelt in spring 2009 and were left in place for five consecutive growing seasons (until end of 2013). They were, however, temporarily dismantled every fall (late October after first snowfall) to protect them from avalanches. They were re-established in the same location every spring immediately after snowmelt. Each size-selective exclosure setup consisted of five plots (2 x 3 m) that progressively excluded aboveground vertebrates and invertebrates from large to small. The plots are labelled according to the guilds that had access to them “L/M/S/I”, “M/S/I”, “S/I”, “I”, “None”; L = large mammals, M = medium mammals, S = small mammals, I = invertebrates, None = no animals had access. As we only had permission to have the experimental setup in place for five consecutive growing seasons, the experiment had to be completely dismantled in the late fall of 2013 and all material removed from the SNP.
Our exclosure design was aimed at excluding mammalian herbivores, but naturally also excluded the few medium and small mammalian predators, as well as the entire aboveground invertebrate food web. A total of 26 large to small mammal species can be found in the SNP, but large apex predators are missing (wolf, bear, lynx) . Reptiles, amphibians and birds are scarce to absent in the subalpine grasslands under study. Only two reptile species occur in the park and they are confined to rocky areas that warm up enough for them to survive. One frog species spawns in an isolated pond far from our grasslands. Only three bird species occasionally feed on the subalpine grasslands. Using game cameras (Moultrie 6MP Game Spy I-60 Infrared Digital Game Camera, Moultrie Feeders, Alabaster, AL, USA), we did observe that the medium- and small-sized mammals (marmot/hares and mice) were not afraid to enter the fences and feed on their designated plots. We never spotted reptiles, amphibians or birds on camera. We distinguished between 59 higher aboveground-dwelling invertebrate taxa that our size-selective exclosures excluded (see also methods for aboveground-dwelling invertebrates below).
The “L/M/S/I” plot (not fenced) was located at least 5 m from the 2.1 m tall and 7 x 9 m large main electrical fence that enclosed the other four plots. The bottom wire of this fence was mounted at 0.5 m height and was not electrified to enable safe access for medium and small mammals, while fencing out the large ones. Within each main fence, we randomly established four 2 x 3 m plots separated by 1-m wide walkways from one another and from the main fence line: 1) the “M/S/I” plots were unfenced, allowing access to all but the large mammals; 2) the “S/I” plots (10 x 10 cm electrical mesh fence) excluded all medium-sized mammals. Note that the bottom 10 cm of this fence remained non-electrified to enable safe access for small mammals; 3) the “I” plots (2 x 2 cm metal mesh fence) excluded all mammals. We double-folded the mesh at the bottom 50 cm to reduce the mesh size to smaller than 1 x 1 cm openings; and 4) the “None” plots were surrounded by a 1 m tall mosquito net (1.5 x 2 mm) to exclude all animals. The top of the plot was covered with a mosquito-meshed wooden frame mounted to the corner posts (roof). We treated these plots a few times with biocompatible insecticide (Clean kill original, Eco Belle GmbH, Waldshut-Tiengen, Germany) to remove insects that might have entered during data collection or that hatched from the soil, but amounts were negligible and did not impact soil moisture conditions within these plots.
To assess whether the design of the “None” exclosure (mesh and roof) affected the response variables within the plots and, therefore, influenced the results, we established an additional six “micro-climate control” exclosures (one in each of the six grasslands) (Risch and others 2013, 2015). These exclosures were built as the “None” exclosures but were open at the bottom (20 cm) of the 3-m side of the fence facing away from the prevailing wind direction to allow invertebrates to enter. A 20-cm high and 3-m long strip of metal mesh was used to block access to small mammals. Thus, this construction allowed a comparable micro-climate to the “None” plots, but also a comparable feeding pressure by invertebrates to the “I” plots. We compared various properties within these exclosures against one another to assess if our construction altered the conditions in the “None” plots. We showed that differences in plant (e.g., vegetation height, aboveground biomass) and soil properties (e.g., soil temperature, moisture) found between the “I” and the “None” treatments were not due to the construction of the “None” exclosure, but a function of animal exclusions, although the amount of UV light reaching the plant canopy was significantly reduced (Risch and others 2013).
Aboveground invertebrate sampling
Aboveground invertebrates were sampled with two different methods to capture both ground- and plant-dwelling organisms: 1) we randomly placed two pitfall traps (67 mm in diameter, covered with a roof) filled with 20% propylene glycol in one 1 x 1 m subplot of the 2 x 3 m treatment plots in spring 2013 (May) and emptied them every two weeks until late September 2013 (Vandegehuchte and others 2017b, 2018). A pitfall trap consisted of a plastic cylinder (13 cm depth, 6.75 cm diameter). Within each cylinder we placed a 100 ml plastic vial with outer diameter 6.70 cm and on top of the cylinder we placed a plastic funnel to guide the invertebrates into the vials. Each trap was cover with a cone-shaped and transparent plastic roof to protect the trap from rain (Vandegehuchte and others 2017b, 2018). Note that in the “None” plots only one trap was placed as control to check for effectiveness of the exclosure. 2) We vacuumed all invertebrates from a 60 x 60 cm area on another 1 x 1 m subplot with a suction sampler (Vortis, Burkhard manufacturing CO, Ltd., Rickmansworth, Hertfordshire, UK) every month from June to September 2013 (Vandegehuchte and others 2017b, 2018). For this purpose, we quickly placed a square plastic frame (60 x 60 x 40 cm) with a closable mosquito mesh sleeve attached to the top edge into the plot from the outside. The suction sample was then inserted into through the sleeve and operated for 45 s to collect the invertebrates (Vandegehuchte and others 2017b, 2018).
We sorted the ≈100 000 individuals collected with both methods by hand and identified each individual morphologically to the lowest taxonomic level feasible (59 taxa, including orders, suborders, subfamilies, families; phylum for Mollusca). These taxa belonged to the following feeding types: 19 herbivores, 16 detritivores, 9 predators, 8 mixed feeders, 5 omnivores and 2 non-classified feeders (or not feeding as adults) (Vandegehuchte and others 2017b). We summed the numbers from the two pitfall traps and the suction sampling over the course of the 2013 season to represent the aboveground invertebrate abundance and community composition of a plot. Note: we did not specifically attempt to catch flying invertebrates with e.g., sticky traps, thus a few flying insects may have been missed with our vacuum sampling approach.
Sampling of plant properties
The vascular plant species composition was assessed at peak biomass every summer (July) by estimating the frequency of occurrence of each species with the pin count method in each plot (Frank and McNaughton 1990). A total of 172 taxa occurred within our 90 plots and we calculated plant species richness for each plot separately. We used the 2013 data in this study. Plant quality was assessed every year in July and September; here we use plant quality at the end of the experiment (September 2013). Two 10 x 100 cm wide strips of vegetation per plot were clipped, combined, dried at 65°C, and ground (Pulverisette 16, Fritsch, Idar-Oberstein, Germany) to pass through a 0.5 mm sieve. Twenty randomly selected samples across all treatments were analysed for N (Leco TruSpec Analyser, Leco, St. Joseph, Michigan, USA) (Vandegehuchte and others 2015). Nitrogen concentrations of the other samples were then estimated from models established for the experiment and the entire SNP relating Fourier transform-near infrared reflectance (FT-NIR) spectra to the measured values of N using a multi-purpose FT-NIR spectrometer (Bruker Optics, Fällanden, Switzerland) (Vandegehuchte and others 2015). Root biomass was sampled every fall by collecting five 2.2 cm diameter x 10 cm deep soil samples (Giddings Machine Company, Windsor, CO, USA) per plot (450 samples year-1). The samples were dried at 30 °C and roots were sorted from the sample by hand. We sorted each sample for 1 h which allowed to retrieve over 90% of all roots present in the samples (Risch and others 2013). The roots were then dried at 65 °C for 48 and weighed to the nearest mg. We averaged the values per plot and used the 2013 data only in this study.
Sampling of edaphic communities
In 2009, 2010, and 2011 we collected three composited soil samples (5 cm diameter x 10 cm depth; AMS Samplers, American Falls, ID, USA) and assessed bacterial community structure using T-RFLP profiling (Liu and others 1997; Blackwood and others 2003; Hodel and others 2014). We detected a total of 89 operational taxonomic units (OTUs). These values are in accordance with other studies reporting OTU richness (Wirthner and others 2011; Zumsteg and others 2012; Meola and others 2014) using T-RFLP profiling, a method that detects the most abundant, and thus likely, the most relevant, taxa. We averaged the data over the three years of collections for our calculations. Microbial biomass carbon (MBC) was determined with the substrate-induced method (Anderson and Domsch 1978) every fall (September) between 2009 and 2013 by collecting three mineral soil samples (5 cm diameter × 10 cm mineral soil core, AMS Samplers, American Falls, ID, USA). The three samples were combined (90 samples for each sampling year), immediately put on ice, taken to the laboratory, passed through a 2-mm sieve and stored at 4°C. Again, we only used the 2013 data in this study.
Soil samples (5 cm diameter x 10 cm depth) to extract soil arthropods were collected in June, July, and August 2011 with a soil corer lined with a plastic sleeve to ensure an undisturbed sample (total of 270 samples). The plastic line core was immediately sealed on both ends using cling film and put into a cooler. All plots were sampled within three days and the extraction of arthropods started the evening of the sampling day using a high-gradient Tullgren funnel apparatus (Crossley and Blair 1991; Vandegehuchte and others 2015). Samples were kept in the extractor for four days and the soil arthropods were collected in 95% ethanol. All individuals were counted and each individual was identified morphologically to the lowest level feasible [76 taxa, including orders, suborders, subfamilies, families (Protura, Thysanoptera, Aphidina, Psylina, Coleoptera, Brachycera, Nematocera, Auchenorryncha, Heteroptera, Formicidae); sub-phylum for Myriapoda, for Acari and Collembola also including morpho-species). Note that we also included larval stages (nine of the 76 taxa) (Vandegehuchte and others 2015). All data were summed over the season. A detailed species list for mites and collembolans is published (Vandegehuchte and others 2017a) [https://doi.org/10.1371/journal.pone.0118679.s001]. Earthworms are rare in the SNP and therefore were not included. We collected eight random 2.2 cm diameter x 10 cm deep soil cores from each plot in September 2013 to determine the soil nematode community composition. The samples were mixed and the nematodes were extracted from 100 ml of fresh soil using Oostenbrink elutriators (Oostenbrink 1960). All nematodes in a 1 ml of the 10 ml extract were counted, a minimum of 150 individuals sample-1 were identified to genus or family level using (Bongers 1988), the numbers of all nematodes were extrapolated to the entire sample and expressed for a 100 g dry sample. In total we identified 63 genus or family levels (Vandegehuchte and others 2015). The list of all the nematodes found is published (Vandegehuchte and others 2015) [http://www.oikosjournal.org/appendix/oik-03341] or DOI: [doi: 10.1111/oik.03341].
We are aware that sampling soil microbes from 2009 to 2011 and soil arthropods in 2011 was not ideal, but we are positive that this does not bias the results. Most of the parameters measured in our experiment either already showed a treatment response after the first growing season (e.g., plant biomass) or did not respond over the entire time experiment (e.g., microbial biomass C). The microbial community composition (2009 – 2011) was highly influenced by inter-annual differences in temperature and precipitation, but did not differ between treatments or vegetation types (Hodel and others 2014). We therefore felt comfortable using the 2009 through 2011 data for describing the soil microbial community in our experimental treatments. Similarly, we are positive that our soil arthropod data are representative. We did assess soil arthropods in August 2012 and found no differences to the August 2011 data. However, we did not feel comfortable combining the 2011 June, July, August data with only August data for 2012 for our analyses.
Sampling of soil properties
We collected three soil samples (5 cm diameter x 10 cm depth) in each plot in September 2013 after removing the vegetation. First, we collected the top layer of mineral soil rich in organic matter, the surface organic layer or rhizosphere, typically 1 to 3 cm in depth with a soil corer (AMS Samples, American Falls, Idaho, USA). Second, we collected a 10 cm mineral soil core beneath this surface layer. The cores for each layer were composited, dried at 65 °C for 48 h and fine-ground to pass a 0.5 mm screen. We then analysed all samples for total C using a Leco TruSpec Analyser (Leco, St. Joseph, Michigan, USA). Mineral soil pH was measured potentiometrically in 1:2 soil:CaCl2 solution with an equilibration time of 30 min.
Soil net N mineralisation was assessed during the 2013 growing season (Risch and others 2015). For this purpose, we randomly collected a 5 cm diameter x10 cm deep soil sample with a soil corer (AMS Samples, American Falls, Idaho, USA) after clipping the vegetation in June 2013. After weighing and sieving (4 mm mesh) the soil, we extracted a 20 g subsample in 1 mol l-1 KCl for 1.5 h on an end-over-end shaker and thereafter filtered it through ashless folded filter paper (DF 5895 150, ALBET LabScience, Hahnenmühle FineArt GmbH, Dassel, Germany). From these filtrates NO3- concentrations were measured colorimetrically (Norman and Stucki 1981) and NH4+with flow injection analysis (FIAS 300, Perkin Elmer, Waltham Massachusetts, USA) (Risch and others 2015). We dried the rest of the sample 105 °C to constant mass to determine fine,fraction bulk density. A second soil sample was collected within each plot in June 2013 with a corer lined with a 5 x 13 cm aluminium cylinder. The corer was driven 11.5 cm deep into the soil so that the top 1.5 cm of the cylinder remained empty. Into this space we placed a polyester bag (250 µm) filled an ion-exchanger resin to capture the incoming N. The bag was filled with a 1:1 mixture of acidic and alkaline exchanger resin (ion-exchanger I KA/ion exchanger IIIAA, Merck AG, Darmstadt, Germany). We then removed 1.5 cm soil at the bottom of the cylinder and placed a second resin exchanger bag into this space to capture the N leached from the soil column. To assure that the exchange resin was saturated with H+ and Cl- prior to filling the bags, the mixture was stirred with 1.2 ml l-1 HCl for 1 h and then rinsed with demineralized water until the electrical conductivity of the water reached 5 µm cm-1. The cylinder with the resin bags in place was reinserted into the soil with the top flush to the soil surface and incubated for three months. We recollected the cylinders in September 2013. Each resin bag and 20 g of sieved soil (4 mm mesh) from each cylinder were then separately extracted with KCl and NO3- and NH4+ concentrations were measured. Nitrate and NH4+ concentrations of all samples were then converted to a content basis by multiplying their values with fine fraction bulk density. Net N mineralisation was thereafter calculated as the difference between the N content of the samples collected at the end of the three-month incubation (including the N extracted from the bottom resin bag) and the N content at the beginning of the incubation (Risch and others 2015).
Soil CO2 emissions were measured every two weeks between 0900 and 1700 hrs from early May through late September 2013 with a PP-Systems SRC-1 soil respiration chamber (15 cm high, 10 cm diameter; closed circuit) attached to a PP-Systems EGM-4 infrared gas analyser (PP-Systems, Amesbury, MA, USA) on two locations per plot (Risch and others 2013). The chamber was placed on randomly placed, permanently installed PVC collars (10 cm diameter) driven 5 cm into the soil at the beginning of the study (Risch and others 2013). Freshly germinated plants growing within the collars were removed prior to each measurement to avoid measuring plant respiration or photosynthesis. The two measurements collected per plot and sampling date were averaged.
Soil moisture (with time domain reflectometry; Field-Scout TDR-100, Spectrum Technologies, Plainfield, Illionois, USA) and temperature (with a waterproof digital pocket thermometer; Barnstead International, Dubuque, Iowa, USA) were measured at five random locations per plot every two weeks during the growing seasons during the experiment for the 0 to 10 cm depth (Risch and others 2013, 2015). As soil moisture and soil temperature were highly negatively correlated (Risch and others 2013), we only used soil moisture for this study. We used plot-level averages of all values available to capture soil moisture variability during the five years of the experiment. The results remained unchanged when we only used soil moisture from the 2013 growing season.
Numeral calculations and statistical analyses
Ecosystem coupling. We conducted principal component analyses (PCAs; unscaled) at the complete dataset level using the abundances of each taxonomical entity to describe each of the five different communities used in this study: aboveground-dwelling invertebrates, vascular plants, soil microorganisms, soil arthropods and soil nematodes. We retained the first two components (PCA axis 1 and PCA axis 2) of each analysis as we found them to adequately represent the temporal and spatial variability of our 90 treatment plots in previous studies55,67. Together they explained a total of 71.70% of the variation for aboveground invertebrates, 44.36% for plants, 44.85% for soil microorganisms, 61.85% for soil arthropods and 77.19% for soil nematodes. In addition, we used soil pH and soil organic C content as a proxy for soil chemical properties, soil bulk density as a proxy for soil physical properties and soil moisture (negatively correlated with soil temperature) as a proxy for soil micro-climatic conditions for an overall total of fourteen constituents.
We calculated ecosystem coupling9 for each exclosure treatment within each vegetation type (i.e., 2  5 treatment combinations in total) as an integrated measure of pairwise ecological interactions between ecosystem constituents representing ecological communities and the soil abiotic environment. These ecological interactions are defined by non-parametric Spearman rank correlation analyses between two constituents, excluding interactions involving two abiotic constituents (e.g., soil pH vs. soil moisture) and interactions between the first (PC1) and second (PC2) component of each community type, as these are orthogonal by definition. Interactions between abiotic constituents were excluded from the analyses because the focus of our study was on communities and how they interact with one another and their surrounding environment; therefore, including abiotic-abiotic interactions was not of interest here. Given that the effectiveness of our experimental design resulted in that no community composition data of aboveground-dwelling invertebrates was available for the “None” plots (all animals excluded), only thirteen instead of fourteen constituents were included in the ecosystem coupling calculations for this treatment. The complete absence of aboveground invertebrates represents the most extreme case of disturbance between aboveground animal communities and the rest of the ecosystem constituents. This may have resulted in a slight overestimation of ecosystem coupling for these plots.
	Average ecosystem coupling was calculated as follows:
Ecosystem coupling=
where Xi is the absolute Coupling was calculated value of the Spearman’s rho coefficient of the ith correlation for each treatment within each vegetation type (i.e., based on nine replicates each), considering and n is the number of pairwise comparisons considered (n = a total of 80; interactions (56 in the case of the “None” treatment). We considered a total of 40 biotic-biotic interactions (i.e., concerning two community-level principal components such as plants and microbes; 24 in the case of the “None” treatment) and 40 abiotic-biotic (i.e., concerning one community-level principal component and one abiotic factor, e.g., plant community and soil properties; 32 in the case of the “None” treatment).	Coupling was calculated for each treatment within each vegetation type (i.e., based on nine replicates each), considering a total of 80 interactions (56 in the case of the “None” treatment). We considered a total of 40 biotic-biotic interactions (i.e., concerning two community-level principal components such as plants and microbes; 24 in the case of the “None” treatment) and 40 abiotic-biotic (i.e., concerning one community-level principal component and one abiotic factor, e.g., plant community and soil properties; 32 in the case of the “None” treatment).
To establish whether constituents were significantly and positively coupled within treatments (i.e., the average of their correlation coefficients were greater than in a null model where correlation only happens by chance), we calculated one-tailed p-values based on permutation tests with 999 permutations.

We considered six ecosystem functions and process rates commonly used to assess ecosystem functioning (Meyer and others 2015; Manning and others 2018). Plant N content represents a measure of forage quality, while plant richness has been shown to stabilise biomass production, thus allowing the system to respond to changes in herbivory. Soil net N mineralisation, soil respiration, root biomass, and microbial biomass represent fluxes or stocks of energy. For all functions and processes higher values represent higher functioning (Manning and others 2018). All these variables were measured in the last year of the experiment (2013). We then quantified ecosystem multifunctionality using the multiple threshold approach (Byrnes and others 2014; Manning and others 2018), which considers the number of functions that are above a certain threshold, over a series of threshold values (typically 10-99%) that are defined based on the maximum value of each function. We weighted all our functions equally for these calculations (Manning and others 2018). The number of functions in a plot with values higher than a given threshold value for the respective function is summed up. The sum represents ecosystem multifunctionality for that plot. Given that choosing any particular threshold as a measure of ecosystem multifunctionality is arbitrary, we calculated the average of thresholds from 10-90% (in 10% intervals) as a more integrated representation of ecosystem multifunctionality.
We used Pearson correlations to explore the relationships between ecosystem coupling (all interactions, biotic-biotic interactions, abiotic-biotic interactions involving above- and belowground constituents, and all interactions, biotic-biotic interactions, abiotic-biotic interactions involving belowground constituents only) and ecosystem multifunctionality by calculating the slopes of all relationships between ecosystem coupling and multifunctionality for all thresholds between 10 and 99%. We also related ecosystem coupling with the average of multifunctionality at thresholds between 30-80% as explained before and considered this correlation as a robust indication of the type of association between these two variables. In addition, we explored the relationships between ecosystem coupling (all interactions, biotic-biotic interactions, abiotic-biotic interactions involving above- and belowground constituents, and all interactions, biotic-biotic interactions, abiotic-biotic interactions involving belowground constituents only) and individual ecosystem functions. The effects of exclosures and vegetation type on individual functions and multifunctionality were evaluated using linear mixed effects models ('lme' function of the nlme package), with exclosure and vegetation type as fixed effects and fence as a random factor. All statistical analyses and numerical calculations were done in R version 3.4.0 (R Core Team 2016).

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- R Core Team. 2016. R: A language and environment for statistical computing. Vienna, Austria: R Foundation for Statistical Computing
- Risch AC, Haynes AG, Busse MD, Filli F, Schütz M. 2013. The response of soil CO2 fluxes to progressively excluding vertebrate and invertebrate herbivores depends on ecosystem type. Ecosystems 16:1192–202.
- Risch AC, Schütz M, Vandegehuchte ML, Van Der Putten WH, Duyts H, Raschein U, Gwiazdowicz DJ, Busse MD, Page-Dumroese DS, Zimmermann S. 2015. Aboveground vertebrate and invertebrate herbivore impact on net N mineralization in subalpine grasslands. Ecology 96:3312–22.
- Schütz M, Risch AC, Achermann G, Thiel-Egenter C, Page-Dumroese DS, Jurgensen MF, Edwards PJ. 2006. Phosphorus translocation by red deer on a subalpine grassland in the Central European Alps. Ecosystems 9:624–633.
- Schütz M, Risch AC, Leuzinger E, Krüsi BO, Achermann G. 2003. Impact of herbivory by red deer (Cervus elaphus L.) on patterns and processes in subalpine grasslands in the Swiss National Park. For Ecol Manage 181:177–88.
- Vandegehuchte ML, van der Putten WH, Duyts H, Schütz M, Risch AC. 2017a. Aboveground mammal and invertebrate exclusions cause consistent changes in soil food webs of two subalpine grassland types, but mechanisms are system-speciﬁc. Oikos 126:212–23.
- Vandegehuchte ML, Raschein U, Schütz M, Gwiazdowicz DJ, Risch AC. 2015. Indirect short- and long-term effects of aboveground invertebrate and vertebrate herbivores on soil microarthropod communities. PLoS One 10:e0118679.
- Vandegehuchte ML, Schütz M, de Schaetzen F, Risch AC. 2017b. Mammal-induced trophic cascades in invertebrate food webs are modulated by grazing intensity in subalpine grassland. J Anim Ecol 86:1434–46.
- Vandegehuchte ML, Trivellone V, Schütz M, Firn J, de Schaetzen F, Risch AC. 2018. Mammalian herbivores affect leafhoppers associated with specific plant functional types at different timescales. Funct Ecol 32:545–55.
- Wirthner S, Frey B, Busse MD, Schütz M, Risch AC. 2011. Effects of wild boar (Sus scrofa L.) rooting on the bacterial community structure in mixed-hardwood forest soils in Switzerland. Eur J Soil Biol 47:296–302. http://dx.doi.org/10.1016/j.ejsobi.2011.07.003
- Zumsteg A, Luster J, Göransson H, Smittenberg RH, Brunner I, Bernasconi SM, Zeyer J, Frey B. 2012. Bacterial, Archaeal and Fungal Succession in the Forefield of a Receding Glacier. Microb Ecol 63:552–64. https://doi.org/10.1007/s00248-011-9991-8

ecosystem-coupling

display_name: ecosystem-coupling

ecosystem-functions

display_name: ecosystem-functions

ecosystem-multifunctionality

display_name: ecosystem-multifunctionality

exclusion

display_name: exclusion

grassland

display_name: grassland

grazing

display_name: grazing

invertebrates

display_name: invertebrates

subalpine

display_name: subalpine

swiss-national-park

display_name: swiss-national-park

vertebrates

display_name: vertebrates

educ

en: Education, culture and sport

envidat

en: EnviDat is the WSL data portal providing unified and managed access to environmental monitoring and research data. The portal has the capability to host and publish data sets. Through the sharing of unique and high-value data sets worldwide, EnviDat fosters research and contributes to advancing environmental science.

en: Ecosystem coupling and multifunctionality - exclosure experiment


maintainer: Jana von Freyberg

en: solutes stable-water-isotopes snowmelt-lysimeter snowmelt-rate snowpack-outflow

state: active

type: dataset

en: The dataset contains volumes, solutes and stable water isotopes of snowpack outflow at daily resolution. 10-minute data are provided of the measurements taken by three snowmelt lysimeter systems at spatially-distributed locations.

en:
This data contain volumes, solutes and isotopes of snowpack outflow measured by a snowmelt lysimeter system at three locations in the southern Alp catchment, situated Central Switzerland. The river Alp is a snow-dominated catchment situated in Central Switzerland characterized by an elevation range from 840 to 1898 m a.s.l.
The dataset provides solutes (major anions and cations, trace metals) and stable water isotopes and water fluxes (snowpack outflow volumes) at daily intervals from several sampling locations. Additionally, the data measured by the snowmelt lysimeter system are provided in 10-minute resolution.

snowmelt-lysimeter

display_name: snowmelt-lysimeter

snowmelt-rate

display_name: snowmelt-rate

snowpack-outflow

display_name: snowpack-outflow

solutes

display_name: solutes

stable-water-isotopes

display_name: stable-water-isotopes

educ

fr: Éducation, culture et sport

de: Bildung, Kultur und Sport

it: Istruzione, cultura e sport

fr: EnviDat: le portail de données environnementales

de: EnviDat: das Umwelt-Datenportal

en: EnviDat: the Environmental Data Portal

it: EnviDat: il portale dei dati ambientali

fr: EnviDat est le portail de données du WSL; il offre un accès uniformisé et réglementé aux données de surveillance et de recherche en matière d’environnement. Ce portail est capable d’héberger et de publier des ensembles de données précieuses et uniques, qui sont partagées aux quatre coins de la planète. Ce faisant, EnviDat favorise la recherche et contribue à l’avancement des sciences environnementales.

de: EnviDat ist das Datenportal der WSL für einen einheitlichen und verwalteten Zugang zu Umweltdaten aus Monitoring- und Forschungsprojekten. Auf dem Portal können Datensätze gehostet und veröffentlicht werden. Durch die weltweite Bereitstellung einzigartiger und wertvoller Datensätze leistet EnviDat einen Beitrag zur Forschung im Bereich der Umweltwissenschaften.

it: EnviDat è il portale dati del WSL che fornisce un accesso unificato e gestito ai dati di monitoraggio ambientale e di ricerca. Il portale ha la capacità di ospitare e pubblicare serie di dati preziosi e unici che vengono condivisi in tutto il mondo. Così facendo, EnviDat promuove la ricerca e contribuisce al progresso della scienza ambientale.

political_level: other

wsl

capacity: public

type: organization

approval_status: approved

key: guid

key: uri

value: Envidat Harvester

en: Daily data of the volumes, solutes and isotopes in snowpack outflow measured at three locations in the southern Alp catchment


maintainer: Athanasios Nenes

en: sigma-w supersaturation raclets cloud-droplets wolfgangpass cloud-modeling

en: Predicted cloud droplet numbers (kappa=0.25) Weissfluhjoch

media_type: text/csv

en:
In-situ measurements of aerosol size distributions along with updraft velocities collected from a wind Doppler Lidar are coupled with a state-of-the art droplet activation parameterization (Morales and Nenes, 2014) to determine the cloud droplet number and maximum in-cloud supersaturation that can potentially form over the Davos Wolfgang station between February 24 and March 8 2019. For this, as a sensitivity test, a hygroscopicity parameter of 0.25 is assumed, which is the upper limit determined from the analysis carried out for Weissfluhjoch (https://www.envidat.ch/dataset/ccn-hygroscopicity-predicted-cloud-droplet-numbers-weissfluhjoch). The data file contains:

* Date and time (DD/MM/YYYY hh:mm:ss) in UTC
* The wind-lidar derived standard deviation of updraft velocities (sigma-w) in m/s, used to apply the parameterization employing a single characteristic velocity (Morales and Nenes, 2010)
* The maximum predicted cloud parcel supersaturation (Smax%)
* The total in-situ aerosol number concentration (Naer) in cmˉ³, measured by the SMPS
* The predicted cloud droplet numbers (Nd) in cmˉ³
* The activation fraction (%)

format: CSV

mimetype: text/csv

en: Predicted cloud droplet numbers (kappa=0.1) Davos Wolfgang

en:
In-situ measurements of aerosol size distributions along with updraft velocities collected from a wind Doppler Lidar are coupled with a state-of-the art droplet activation parameterization (Morales and Nenes, 2014) to determine the cloud droplet number and maximum in-cloud supersaturation that can potentially form over the Davos Wolfgang station between February 24 and March 8 2019. For this, as a sensitivity test, a hygroscopicity parameter of 0.1 is assumed, which is the lower limit determined from the analysis carried out for Weissfluhjoch (https://www.envidat.ch/dataset/ccn-hygroscopicity-predicted-cloud-droplet-numbers-weissfluhjoch). The data file contains:

* Date and time (DD/MM/YYYY hh:mm:ss) in UTC
* The wind-lidar derived standard deviation of updraft velocities (sigma-w) in m/s, used to apply the parameterization employing a single characteristic velocity (Morales and Nenes, 2010)
* The maximum predicted cloud parcel supersaturation (Smax%)
* The total in-situ aerosol number concentration (Naer) in cmˉ³, measured by the SMPS
* The predicted cloud droplet numbers (Nd) in cmˉ³
* The activation fraction (%)

en: Cloud droplet properties were predicted between February 24 and March 8 2019 for the measurement site Davos Wolfgang (1630 m a.s.l., LON: 9.853594, LAT: 46.835577). Droplet calculations are carried out with the physically based aerosol activation parameterization of Morales and Nenes (2014), employing the “characteristic velocity” approach of Morales and Nenes (2010). Aerosol size distribution observations required to predict the cloud droplet numbers and maximum in-cloud supersaturation are obtained from a Scanning Mobility Particle Size Spectrometer (SMPS) instrument deployed at Davos Wolfgang (https://www.envidat.ch/dataset/aerosol-data-davos-wolfgang). The required vertical velocity measurements are derived from the wind Doppler Lidar (https://www.envidat.ch/dataset/lidar-wind-profiler-data) deployed at the same station and are extracted from the first bin of the instrument, being 200 m above ground level. The hygroscopic properties of the particles measured at Davos Wolfgang could not be estimated, owing to a lack of concurrent CCN measurements. As a sensitivity test, droplet calculations are performed assuming two different values of the aerosol hygroscopicity parameter, 0.1 and 0.25, based on the analysis carried out for Weissfluhjoch. Additional information can be found in Section 2.3 [here](https://acp.copernicus.org/preprints/acp-2020-1036/).

cloud-droplets

display_name: cloud-droplets

cloud-modeling

display_name: cloud-modeling

raclets

display_name: raclets

sigma-w

display_name: sigma-w

supersaturation

display_name: supersaturation

wolfgangpass

display_name: wolfgangpass

en: Predicted cloud droplet numbers Davos Wolfgang


maintainer: Marcus Schaub

en: diameter-at-breast-height dbh tree-species swiss-forest-lab tree drought forest irrigation scots-pine lwf crown-condition

en: Site photo 3

format: SERVICE

en: Shapefile	of measurement location

en: Swiss Forest Health Inventory

media_type: text/html

format: HTML

mimetype: text/html

en: Dataset

en: Site photo 1

en: Site photo 2

en: Long-term Irrigation	Experiment Pfynwald homepage

en: To study the performance of mature Scots pine (_Pinus sylvestris_ L.) under chronic drought conditions in comparison to their immediate physiological response to drought release, a controlled long-term and large-scale irrigation experiment has been set up in 2003. The experiment is located in a xeric mature Scots pine forest in the Pfynwald (46° 18' N, 7° 36' E, 615 m a.s.l.) in one of the driest inner-Alpine valleys of the European Alps, the Valais (mean annual temperature: 9.2°C, annual precipitation sum: 657 mm, both 1961-1990). Tree age is on average 100 years, the top height is 10.8 m and the stand density is 730 stems ha-1 with a basal area of 27.3 m2 ha-1. The forest is described as _Erico Pinetum sylvestris_ and the soil is a shallow pararendzina characterized by low water retention. The experimental site (1.2 ha; 800 trees) is split up into eight plots of 1'000 m2 each. During April-October, irrigation is applied on four randomly selected plots with sprinklers of 1 m height at night using water from an adjacent water channel. The amount of irrigation corresponds to a supplementary rainfall of 700 mm year-1. Trees in the other four plots grow under naturally dry conditions. Soil moisture has been monitored since the beginning of the project at 3 soil depths (10, 20 and 60 cm). The crown condition of each tree is being assessed each year since 2003. Tree measurement data such as diameter at breast height, tree height, and social status were assessed in 2002, 2009 and 2014. The duration of the irrigation experiment is planned for 20 years.

crown-condition

display_name: crown-condition

dbh

display_name: dbh

diameter-at-breast-height

display_name: diameter-at-breast-height

drought

display_name: drought

forest

display_name: forest

irrigation

display_name: irrigation

lwf

display_name: lwf

scots-pine

display_name: scots-pine

swiss-forest-lab

display_name: swiss-forest-lab

tree

display_name: tree

tree-species

display_name: tree-species

en: Tree measurements 2002-2016 from the long-term irrigation experiment Pfynwald, Switzerland


en: social-acceptance optimization photovoltaic ecosystem-services marxan energy wind-power spatial-planning landscape

media_type: application/zip

en:
This is the compressed Shapefile (ESRI) from the MARXAN optimization with 2216 rows (objects) and 18 columns (variables) with sf (simple feature) and data frame classes.
Column description:
X01 = Planning_units (PUs)
X02 = incl.OS. number of times pu was selected in MARXAN when optimized for ecological costs
X03 = incl.OS. number of times pu was selected in MARXAN when optimized for social costs
X04 = incl.OS. ecological costs (ECUess) of pu when realizing the total energy (normalized)
X05 = incl.OS. social costs (ECUsoc) of pu when realizing the total energy (normalized)
X06 = incl.OS. total energy (MWh/a)
X07 = incl.OS. energy from roof-mounted PV infrastructure (MWh/a)
X08 = incl.OS. energy from wind turbines (MWh/a)
X09 = incl.OS. energy from ground-mounted PV infrastructure (MWh/a)

X10 = excl.OS. number of times pu was selected in MARXAN when optimized for ecological costs
X11 = excl.OS. number of times pu was selected in MARXAN when optimized for social costs
X12 = excl.OS. ecological costs (ECUess) of pu when realizing the total energy (normalized)
X13 = excl.OS. social costs (ECUsoc) of pu when realizing the total energy (normalized)
X14 = excl.OS. total energy (MWh/a)
X15 = excl.OS. energy from roof-mounted PV infrastructure (MWh/a)
X16 = excl.OS. energy from wind turbines (MWh/a)
X17 = excl.OS. energy from ground-mounted PV infrastructure (MWh/a)
geometry = Simple feature XY geometry (SFC_POINT) each representing the center of a 4x4km planning unit (PU) in EPSG 21781 (CRS CH1903 / LV03).

format: ZIP

en:
This study examines the siting scenarios for renewable energy infrastructure (REI) in Switzerland, incorporating the external costs of ecosystem services and, innovatively, social preferences. This approach challenges the prevalent techno-economic siting paradigm, which often overlooks these externalities. To minimize the external costs of the scenarios while maximizing energy yield, Marxan, an optimization software, was employed.
MARXAN was run for 2 versions: a) without ground-mounted open space PV infrastructure (excl. OS) and b) with ground-mounted open space PV infrastructure (incl. OS). In each version optimization was done using ecological costs (ECUess) or social costs (ECUsoc) in a regular grid of 4x4km (planning unit).

File:
PU_data.shp: compressed Shapefile (ESRI) from the MARXAN optimization with 2216 rows (objects) and 18 columns (variables) with sf (simple feature) and data frame classes.

Headers are described below:
X01 = Planning_units (PUs)

X02 = incl.OS. number of times pu was selected in MARXAN when optimized for ecological costs
X03 = incl.OS. number of times pu was selected in MARXAN when optimized for social costs
X04 = incl.OS. ecological costs (ECUess) of pu when realizing the total energy (normalized)
X05 = incl.OS. social costs (ECUsoc) of pu when realizing the total energy (normalized)
X06 = incl.OS. total energy (MWh/a)
X07 = incl.OS. energy from roof-mounted PV infrastructure (MWh/a)
X08 = incl.OS. energy from wind turbines (MWh/a)
X09 = incl.OS. energy from ground-mounted PV infrastructure (MWh/a)

X10 = excl.OS. number of times pu was selected in MARXAN when optimized for ecological costs
X11 = excl.OS. number of times pu was selected in MARXAN when optimized for social costs
X12 = excl.OS. ecological costs (ECUess) of pu when realizing the total energy (normalized)
X13 = excl.OS. social costs (ECUsoc) of pu when realizing the total energy (normalized)
X14 = excl.OS. total energy (MWh/a)
X15 = excl.OS. energy from roof-mounted PV infrastructure (MWh/a)
X16 = excl.OS. energy from wind turbines (MWh/a)
X17 = excl.OS. energy from ground-mounted PV infrastructure (MWh/a)

geometry = Simple feature XY geometry (SFC_POINT) each representing the center of a 4x4km planning unit (PU) in EPSG 21781 (CRS CH1903 / LV03).

For MARXAN optimizations (for meaning of standard files in MARXAN see https://scholar.google.ch/scholar_url?url=https://courses.washington.edu/cfr590/software/Marxan1810/marxan_manual_1_8_2.pdf&hl=de&sa=X&ei=NAWHZI6JJ8PFmAGMo47oDQ&scisig=AGlGAw-DBrsV4kUzkR6GkN1jtG66&oi=scholarr), pu_data.shp was used to generate MARXAN files input.dat, pu.dat, puvfeat.dat, spec.dat.
Coordinate Reference System: CH1903 / LV03. EPSG: 21781.

ecosystem-services

display_name: ecosystem-services

energy

display_name: energy

landscape

display_name: landscape

marxan

display_name: marxan

optimization

display_name: optimization

photovoltaic

display_name: photovoltaic

social-acceptance

display_name: social-acceptance

spatial-planning

display_name: spatial-planning

wind-power

display_name: wind-power

educ

en: Education, culture and sport

envidat

en: EnviDat: the Environmental Data Portal

en: EnviDat is the WSL data portal providing unified and managed access to environmental monitoring and research data. The portal has the capability to host and publish data sets. Through the sharing of unique and high-value data sets worldwide, EnviDat fosters research and contributes to advancing environmental science.

en: Optimizing renewable energy siting in the Swiss landscape


maintainer: EnviDat Support

en: roads gis jupyter geospatial density buffer

en: Road data from OSM for the surroundings of Menzberg, Switzerland

mimetype: application/zip

en: Plots

en: Random plot locations

en: Road Density Calculation (Python Notebook) online

en:
This dataset contains the road and plot data used for the geospatial analysis example showcased in "Fostering Open Science at WSL with the EnviDat Environmental Data Portal", a contribution to the 5th Open Source Geospatial Research and Education Symposium (OGRS), 2018.
The example uses Jupyter Notebook to calculate road densities in the neighbourhood of sample plot locations with Python. Road data were extracted from OpenStreetMap, while the point data (sample plots) were generated manually.

buffer

display_name: buffer

density

display_name: density

geospatial

display_name: geospatial

gis

display_name: gis

jupyter

display_name: jupyter

roads

display_name: roads

en: Dataset for OGRS 2018 publication


maintainer: Matthias Haeni

en: long-term par meteorology temperature climate-change relative-humidity precipitation meteo swiss-forest-lab wind-speed

en: Data plot of the field station (JUF)

media_type: application/pdf

en: Data overview for air temperature, relative humidity, precipitation, PAR, wind speed.

format: PDF

mimetype: application/pdf

en: Forest station data (JUB)

en: Data file.

en: Data plot of the forest station (JUB)

en: Site photo of the field station (JUF)

en: Site photo of the field station from the side.

mimetype: image/jpeg

en: Site photo of the forest station (JUB)

en: Site photo of the forest station from the side.

en: Site aerial photo (Jussy)

en: Site aerial photo from both Jussy stations. Source: Google. 2016. Site aerial photo from both Jussy stations. Retrieved November 11, 2016, from https://www.google.ch/maps

en: High quality meteorological data are needed for long-term forest ecosystem research, particularly in the light of global change. The long-term data series published here comprises almost 20 years of measurements for two meteorological stations in Jussy in Switzerland where one station is located within a natural broad-leaved forest stand (JUB) with sessile oak (_Quercus petrea_; 90 yrs), aspen (_Populus tremula_; 60 yrs) and European hornbeam (_Carpinus betulus_; 60 yrs) as dominant tree species. A second station is situated in the very vicinity outside of the forest (field station, JUF). The meteorological time series are presented in hourly time resolution of air temperature, relative humidity, precipitation, photosynthetically active radiation (PAR) and wind speed. Jussy is part of the Long-term Forest Ecosystem Research Programme (LWF) established and maintained by the Swiss Federal Research Institute WSL.

climate-change

display_name: climate-change

long-term

display_name: long-term

meteo

display_name: meteo

meteorology

display_name: meteorology

par

display_name: par

precipitation

display_name: precipitation

relative-humidity

display_name: relative-humidity

swiss-forest-lab

display_name: swiss-forest-lab

temperature

display_name: temperature

wind-speed

display_name: wind-speed

en: Jussy, Switzerland: Long-term forest meteorological data from the Long-term Forest Ecosystem Research Programme (LWF), from 1997 onwards


maintainer: Armin Sigmund

en: evaporation antarctica stable-water-isotopes backward-trajectories isotopic-distillation lagrangian-analysis snow-sublimation

en: Model code

en: Model Sublimation and Model Air Parcel (python scripts)

en: Data

en: Modeled data including the data shown in the figures of the related research article

en: Python scripts for data visualization

en: Scripts for generating the figures of the related research article and generating the data files (csv) from the model output

en: Readme

media_type: text/plain

en: Meta data

format: TXT

en:
 Summary

This data set contains Python programming code and modeled data discussed in a related research article. We developed a simple isotope model to study the drivers of the particularly depleted vapour isotopic composition measured on the ship of the Antarctic Circumnavigation Expedition close to the outlet of the Mertz glacier, East Antarctica, in the 6-day period from 27 January 2017 to 1 February 2017. The model considers the stable water isotopologues H2(16O), H2(18O), and HD(16O). It uses data from the ERA5 reanalysis product with a spatial resolution of 0.25° x 0.25° (Hersbach et al., 2018) and 10-day backward trajectories for the location of the ship, published by Thurnherr et al. (2020a). Our data set includes the model code, Python scripts for visualizing the results, and data produced by the model including the results shown in the figures of the related research article. Here, we summarize the most important model characteristics while further details can be found in the readme.txt file and the related research article including its supporting information.

 Main model characteristics

The modeling approach consists of two steps called *Model Sublimation* and *Model Air Parcel*. The former estimates the isotopic compositions of the snow and sublimation flux across the Antarctic Ice Sheet using an Eulerian frame of reference while the latter models the vapour isotopic composition and specific humidity along air parcel trajectories using a Lagrangian frame of reference. The isotope effects of most phase changes are represented by equilibrium fractionation. Only for ocean evaporation, kinetic fractionation is additionally taken into account (original Craig-Gordon formula). For snow sublimation, two assumptions are tested: *Run E* assumes that sublimation is associated with equilibrium fractionation while *Run N* assumes that sublimation occurs without isotopic fractionation.

 Model Sublimation
Model Sublimation uses a simple one-dimensional mass-balance approach in each grid cell, considering snow accumulation due to snowfall and vapour deposition and snow ablation due to sublimation. The snowpack is represented by 100 layers of equal thickness (e.g., 1 cm) and density (350 kg m-3). The isotopic composition of snowfall is parameterized by generalizing a site-specific, empirical relationship between the daily mean air temperature and snowfall isotopic composition. In the case of vapour deposition, Model Sublimation assumes equilibrium fractionation and estimates the isotopic composition of the atmospheric vapour as the average value for two idealized situations: (i) locally sourced vapour which has the same isotopic composition as the sublimation flux; (ii) non-locally sourced vapour in isotopic equilibrium with snowfall. Model Sublimation is run with a time step of 1 h, independently of Model Air Parcel.

 Model Air Parcel
Every hour, an ensemble of trajectories arrives at different heights in the ABL above the ship. For each of these trajectories, we consider an air parcel with a constant volume of 1 x 1 x 1 m3. The air parcels are initialized at the first suitable time when the trajectories are located in the ABL, either over the ice-free ocean in conditions of evaporation or over snow (Antarctic Ice Sheet or sea ice). Subsequently, the masses of the water isotopologues in the air parcels are simulated with a time step of 3 h, considering vapour uptake or removal due to the moisture flux at the snow or liquid ocean surface (only if the parcel is in the ABL) and cloud/precipitation formation (if the saturation specific humidity is reached). Sea ice is taken into account in a very simplified way. We represent the sea ice by grid cells with a sea-ice cover of more than 90% and assume the isotopic composition of the sublimation flux to be identical to that in the nearest grid cell of the Antarctic Ice Sheet. The isotopic composition of the sublimation flux is taken from Model Sublimation whereas the isotopic composition of the vapour deposition flux (over snow) and condensation flux (over ice-free ocean) is simulated assuming an isotopic equilibrium with the air parcel. Isotope effects of cloud/precipitation formation are represented using the classic Rayleigh distillation model with equilibrium fractionation, where the cloud water is assumed to precipitate immediately after formation.

 References

Hersbach, H., Bell, B., Berrisford, P., Biavati, G., Horanyi, A., Munoz Sabater, J.,... others (2018). *ERA5 hourly data on single levels from 1979 to present. Copernicus Climate Change Service (C3S) Climate Data Store (CDS)*. doi: 10.24381/cds.bd0915c6

Thurnherr, I., Wernli, H., & Aemisegger, F. (2020a). *10-day backward trajectories from ECMWF analysis data along the ship track of the Antarctic Circumnavigation Expedition in austral summer 2016/2017*. Zenodo. doi: 10.5281/zenodo.4031705

antarctica

display_name: antarctica

backward-trajectories

display_name: backward-trajectories

evaporation

display_name: evaporation

isotopic-distillation

display_name: isotopic-distillation

lagrangian-analysis

display_name: lagrangian-analysis

snow-sublimation

display_name: snow-sublimation

stable-water-isotopes

display_name: stable-water-isotopes

educ

en: Education, culture and sport

envidat

en: EnviDat: the Environmental Data Portal

en: EnviDat is the WSL data portal providing unified and managed access to environmental monitoring and research data. The portal has the capability to host and publish data sets. Through the sharing of unique and high-value data sets worldwide, EnviDat fosters research and contributes to advancing environmental science.

en: Modeled Isotopic Composition of Water Vapour Along Air Parcel Trajectories in the Antarctic


radiosondes

maintainer: Jörg Wieder

title_for_slug: radiosondes

en: raclets wind-direction relative-humidity pressure temperature radiosonde wind holoballoon

en: Radiosond Template Files

en:
The "template" files contain standard parameters, i.e. ambient pressure (mbar and Pa), temperature (°C), relative humidity (%), wind speed (m s‾ ¹) and wind direction (true deg), as well as additional information, e.g. time (UTC and local time), dew point (°C), mixing ration (g/kg), density (kg m³), altitude (m MSL), height above ground (ft and m AGL), and geographic coordinates (longitude, latitude). If the wind speed was zero, no wind direction is given.
The zip files contain ascents on the following days:
February 10 (morning and afternoon),
February 11 (morning),
February 16 (afternoon),
February 22 (morning),
March 1 (morning),
March 5 (morning),
March 11 (morning),
March 12 (morning),
March 13 (morning),
March 14 (morning),
March 17 (afternoon), and
March 18.
The data was managed with Windsond Receiver Application Demo Version 2.92, (C) Sparv Embedded AB 2016.

en: Radiosond Sounding Files

en:
The "soundings" radiosond data contains information on ambient pressure (mbar), temperature (C), relative humidity (%), wind speed (m s‾ ¹) and wind direction (true deg), and are ordered with respect to the height above ground (m AGL) with a step size of 20 m. The first line of each file idicates the geographic location and the start time (in UTC) of the ascent. If the wind speed was zero, no wind direction is given.
The zip files contain ascents on the following days:
February 10 (morning and afternoon),
February 11 (morning),
February 16 (afternoon),
February 22 (morning),
March 1 (morning),
March 5 (morning),
March 11 (morning),
March 12 (morning),
March 13 (morning),
March 14 (morning),
March 17 (afternoon), and
March 18.
The data was managed with Windsond Receiver Application Demo Version 2.92, (C) Sparv Embedded AB 2016.

en: Radiosondes (Windsond, Sparv Embedded AB) were started in Davos Wolfgang (LON: 9.853594, LAT: 46.835577) to report height profiles of pressure, relative humidity and temperature at specific days. In addition to regular launches of radiosondes, sondes were attached to [HoloBalloon](https://www.envidat.ch/group/clouds-in-situ-raclets) to report the ambient conditions of the in-situ measurements. Further profiles of meteorological measures were recorded at [HoloGondel](https://www.envidat.ch/group/clouds-in-situ-raclets) which was installed at the gondola moving between Gotschnaboden and Gotschnagrat at 2285 m a.s.l.

holoballoon

display_name: holoballoon

pressure

display_name: pressure

raclets

display_name: raclets

radiosonde

display_name: radiosonde

wind

display_name: wind

wind-direction

display_name: wind-direction

en: Radiosondes


en: shortwave-radiation longwave-radiation weisfluhjoch davos radiation

en: Processing doc

en: Description of processing code that was ported to R by P. Frei (2016).

en: Parameter file

en: Description of ASRB parameters.

mimetype: text/plain

en: Metadata

en: DATA ACCESS

en: ASRB radiation station at the Weissfluhjoch versuchsfeld.

en:
Incoming and outgoing shortwave and longwave 2 min radiation measurements at the Weissfluhjoch research site, Davos, CH. The experimental site at the Weissfluhjoch (WFJ, 46.83 N, 9.81 E) is located at an altitude of 2540 m in the Swiss Alps near Davos. During the winter months, almost all precipitation falls as snow at this altitude. As a consequence, a continuous seasonal snow cover builds up every winter, with a maximum snow height ranging from 153–366 cm over the period 1934–2012. The measurement site is located in an almost flat part of a southeast oriented slope.

 References
1. Marty, C., Philipona, R., Frohlich, C., Ohmura, A.. Altitude dependence of surface radiation fluxes and cloud forcing in the alps: results from the alpine surface radiation budget network. 2002. Theoretical and Applied Climatology. Volume 72. Issue 3-4. 137-155. http://dx.doi.org/10.1007/s007040200019. 10.1007/s007040200019.
2. Christoph Marty. Surface Radiation, Cloud Forcing and Greenhouse Effect in the Alps. 2000. Institute fuer Klimaforschung ETH. Zuercher Klima-Schriften. Volume 79. http://e-collection.library.ethz.ch/eserv/eth:23491/eth-23491-01.pdf.

davos

display_name: davos

longwave-radiation

display_name: longwave-radiation

radiation

display_name: radiation

shortwave-radiation

display_name: shortwave-radiation

weisfluhjoch

display_name: weisfluhjoch

en: ASRB_WFJVF: Shortwave and longwave radiation measurements (2 min) at the Weissfluhjoch research site, Davos


en: drifting-snow large-eddy-simulation saltation-parameterizations aerodynamic-entrainment snow-saltation splash

en: Code and input files to reproduce the simulations in Melo et al. (2024), as well as the post-processing scripts used in the analysis.

en:
This dataset includes the necessary files to reproduce the results presented in Melo et al. (2024):
- code version used in the simulations (also available at the institutional [GitLab repository](https://gitlabext.wsl.ch/atmospheric-models/les-lsm) - tag 'r2024')
- input files to run the simulations
- post-processing scripts (MATLAB and FORTRAN)
- selected raw output files
- post-processed results (MATLAB data files).
 	**Note:** Direct model outputs are not provided because of their size, but they can be shared upon request. They can be reproduced by running the code. The post-processed results allow you to reproduce all figures in Melo et al. (2024).

aerodynamic-entrainment

display_name: aerodynamic-entrainment

drifting-snow

display_name: drifting-snow

large-eddy-simulation

display_name: large-eddy-simulation

saltation-parameterizations

display_name: saltation-parameterizations

snow-saltation

display_name: snow-saltation

splash

display_name: splash

educ

en: Education, culture and sport

envidat

en: EnviDat: the Environmental Data Portal

en: EnviDat is the WSL data portal providing unified and managed access to environmental monitoring and research data. The portal has the capability to host and publish data sets. Through the sharing of unique and high-value data sets worldwide, EnviDat fosters research and contributes to advancing environmental science.

en: Understanding snow saltation parameterizations - Reproducibility


en: ice-layers snowmicropen weissfluhjoch snow-stratigraphy snow-temperature water-front

en: README

en: WFJ_ICE_LAYERS dataset

en: The WFJ_ICE_LAYERS dataset contains multi-instrument snowpack measurements at high temporal resolution, which enable to monitor the formation of deep ice layers due to preferential water flow, at the Weissfluhjoch research site, Davos, Switzerland. It covers the winter 2016/2017, with a focus on the early melting season. This dataset includes traditional snowpack profiles (weekly resolution, 15/11/2016-29/05/2017), SnowMicroPen penetration resistance profiles (daily resolution, 01/02/2017-19/04/2017), snow temperatures measured at different heights in the snowpack (half-hourly resolution, 01/03/2017-15/04/2017) and the water front height derived from an upward-looking ground penetrating radar (3-hour resolution, 04/03/2017-08/04/2017). The measurements are complemented by initialization files for SNOWPACK model simulations with the ice reservoir parameterization at Weissfluhjoch for the winter 2016/2017.

ice-layers

display_name: ice-layers

snow-stratigraphy

display_name: snow-stratigraphy

snow-temperature

display_name: snow-temperature

snowmicropen

display_name: snowmicropen

water-front

display_name: water-front

weissfluhjoch

display_name: weissfluhjoch

en: WFJ_ICE_LAYERS: Multi-instrument data for monitoring deep ice layer formation in an alpine snowpack


maintainer: Giulia Francesca Azzurra Donati

en: ecosystem-services urban-planning biodiversity ecological-corridors circuit-theory network-analysis

en: Amphibian movement resistance layers and scripts for connectivity modelling - part 2

en: The zipped folder includes the script to create and analyse the networks from Circuitscape outputs. The subfolders “Boundaries”, “Layers”, and “Reference_net” contain files either called by the code or created in the process. The subfolder “Outputs_from_Circuitscape” contains the outputs from Circuitscape including node and branch cumulative currents in a network format.

en: Amphibian movement resistance layers and scripts for connectivity modeling - part 1

en: The zipped folder includes the script to aggregate the resistance layers into squared and hexagonal grids. The subfolders “Boundaries”, “Layers”, “Reference_grids”, and “Resistance_from_SDMs” contain files either called by the code or created in the process.The subfolder “Inputs_for_Circuitscape” contains the inputs to run Circuitscape, after doing some subsequent processing steps in QGIS (listed in the R script) with the files created with the script.

en: This data includes (1) Scripts to aggregate landscape resistance layers into squared and hexagonal grids (i.e., different representations and resolutions), (2) Input resistance layers and focal nodes in .txt format to run in Circuitscape (Python implementation v4.0.5). Circuitscape is a software tool for modeling and analyzing landscape connectivity, which simulates movement of organisms across landscapes by estimating resistance to movement across each point of the landscape. (3) Scripts for the ecological network analysis, and (4) environmental predictors for amphibian whole-life cycle habitats used to describe the local environment for BGI design (i.e. topographic, hydrologic, edaphic, vegetation, land-use derived, movement-ecology related).

biodiversity

display_name: biodiversity

circuit-theory

display_name: circuit-theory

ecological-corridors

display_name: ecological-corridors

ecosystem-services

display_name: ecosystem-services

network-analysis

display_name: network-analysis

urban-planning

display_name: urban-planning

en: Multi-Scale Prioritization framework for Urban Blue-Green Infrastructure Planning to Support Biodiversity: Data & Codes


en: microct intercomparison specific-surface-area snow icecube

en: Grain survey of snow particles

en: The file contains all measurements of the artificial particles for snow types A, B, C, D and E, including the artificial particle mass in the Petri dish.

en: artificial_snow_data set

en: The file contains all micro-CT and IC measurements of the artificial snow data set. The data set was originally produced to investigate the influence of black carbon on SSA measurements.

en: The file contains all data for the plot in figure 1 of the co-located density and SSA measurement at Summit, Greenland in May 2015, conducted by Schneebeli, Proksch, Schmid.

en: IceCube and MicroCT data

en: The file contains all micro-CT and IC measurements for snow types A, B, C, D and E used in our study. All samples were processed (and stored) in a cold laboratory at -15°C.

en: TARTES simulations

en: Jupyter notebook with TARTES simulations.

en: Influence of black carbon on the measurement of the specific surface of snow with the near infrared method - Philipp Baumann

en: These experiments investigate different possible errors by measuring the specific surface area (SSA) of snow with the near infrared method. The influence of black carbon on this measurement was disproved, the theory was therefore validated. It shows that sunlight has no influence on the measurement with the IceCube. Wind drift snow as thin layer on the sample can increase the measured value of the specific surface by a few percent. This could be part of the error, observed in former measurements at SLF. As reference for the measured SSA values with the near infrared method, computer tomography was used to verify the SSA values.

en: Statistics of samples

en: SMP measurement files for snow types A, B, C and D.

en: The specific surface area (SSA) of different snow types were measured with the IceCube instrument and the Scanco Medical microCT 40. In addition, the snow particles created during the preparation of IceCube samples were counted. The difference in SSA between these instruments is explained by the formation of the surface particles. A numerical simulation using TARTES simulation support the observations.

icecube

display_name: icecube

intercomparison

display_name: intercomparison

microct

display_name: microct

snow

display_name: snow

specific-surface-area

display_name: specific-surface-area


en: climate-change phytochemistry herbivory drought-legacy fagus-sylvatica

en:
What are the research data files about:
Raw data on various beech (Fagus sylvatica) leaf traits. Beech leaf chemistry (primary and specialized metabolites), leaf damage measurements from various herbivore feeding guilds and crown health assessment

Which methods were used:
Field surveys on trees that showed either prematured defoliation due to drought-stress or showed not pre-mature defoliation. .

When and where was the data collected/analyzed:
Field data was collected in 2019, 2020
Data was analyzed in 2021-2023

climate-change

display_name: climate-change

drought-legacy

display_name: drought-legacy

fagus-sylvatica

display_name: fagus-sylvatica

herbivory

display_name: herbivory

phytochemistry

display_name: phytochemistry

educ

en: Education, culture and sport

envidat

en: EnviDat: the Environmental Data Portal

en: EnviDat is the WSL data portal providing unified and managed access to environmental monitoring and research data. The portal has the capability to host and publish data sets. Through the sharing of unique and high-value data sets worldwide, EnviDat fosters research and contributes to advancing environmental science.

en: Legacy effects of premature defoliation in response to an extreme drought event modulate phytochemical profiles with subtle consequences for leaf herbivory in European beech


en: longwave-radiation shortwave-radiation outgoing davos incoming

en: Metadata

en: Processing doc

en: Parameter file

en: DATA ACCESS

en:
Incoming and outgoing shortwave and longwave 2 min radiation measurements in Davos Dorf, CH.

 References
1. Marty, C., Philipona, R., Frohlich, C., Ohmura, A.. Altitude dependence of surface radiation fluxes and cloud forcing in the alps: results from the alpine surface radiation budget network. 2002. Theoretical and Applied Climatology. Volume 72. Issue 3-4. 137-155. http://dx.doi.org/10.1007/s007040200019. 10.1007/s007040200019.
2. Christoph Marty. Surface Radiation, Cloud Forcing and Greenhouse Effect in the Alps. 2000. Institute fuer Klimaforschung ETH. Zuercher Klima-Schriften. Volume 79. http://e-collection.library.ethz.ch/eserv/eth:23491/eth-23491-01.pdf.

davos

display_name: davos

incoming

display_name: incoming

longwave-radiation

display_name: longwave-radiation

outgoing

display_name: outgoing

shortwave-radiation

display_name: shortwave-radiation

en: ASRB_DAV: Shortwave and longwave radiation measurements (2 min) in Davos Dorf


en: land-use-simulations spatial-planning land-use-change digital-zoning-plans land-use-transformations

en: Legend simulated maps

en:
Table of Content: 1. General context of the data set "LUSzoning”; 2. Background and aims of the study using the data set LUSzoning; 3. The data set LUSzoning.
1. General context of the data set "LUSzoning".
The data set "LUSzoning" stands for Land-use simulations integrating zoning regulations in Spanish functional urban areas. The data set has been generated as part of the CONCUR research project (https://www.wsl.ch/en/projects/concur.html) led by Dr. Anna M. Hersperger and funded by the Swiss National Science Foundation (ERC TBS Consolidator Grant (ID: BSCGIO 157789) for the period 2016-2021. The CONCUR research project is interdisciplinary and aims to develop a scientific basis for adequately integrating spatial policies (in this case, digital zoning plans) into quantitative land-change modelling approaches at the urban regional level.
2.	Background and aims of the study using the data set “LUSzoning”.
As part of the CONCUR project, a specific task was to integrate planning spatial policies in land-change modelling. Planning can be implemented in modelling using either hard or gradual restrictions. Different studies have addressed the inclusion of spatial planning policies in land-use change modelling. However, the integration of zoning constraints is generally established as hard or Boolean-based restrictions (e.g., whether urban development is allowed or not), while not accounting for the spatial heterogeneity or gradual characteristics within planning zones (e.g., whether planning regulations allow low, medium or high urban density), though these could improve real patterns simulations in urban areas. We assume Spanish General Zoning plans were suitable to explore the integration of planning into land-change modelling as soft constrains because they define land-use intensities in the buildable zoning areas. In light of the above considerations, the overall aim of the study was to model urban land-use changes using a multi-scenario approach that integrates digitized zoning plans for the Functional Urban Areas (FUAs) of Madrid, Barcelona, Valencia, and Zaragoza. The following specific objectives were addressed: i) to analyse the role of planning by defining three future scenarios that integrate digitized zoning plans and one scenario that assumes almost no planning intervention; ii) to introduce zoning constraints that reflect different degrees of urban densities; iii) to generate a transferable spatially-explicit modelling framework to integrate planning into land-use change simulations.
Four future land-use demands scenarios were defined for the FUAs. Storylines were created considering probable development scenarios related to zoning plans, current Spanish legislation and sustainability goals defined along two axes: a high market-oriented vs. high planning-intervention axis, and an axis of short-term economic growth vs. long-term sustainable growth. The sustainable development scenario (S1) is characterized by low gross floor area (GFA) growth that is limited to areas that are currently under development according to zoning plans. The business-as-usual scenario (S2) is characterized by medium GFA growth in the range of on-going trends. The strong development scenario (S3) is characterized by high GFA growth rates. Growth is restricted to buildable areas without urbanization project designated in zoning plans. The unrestricted development scenario (S4) prioritizes a high degree of market liberalization characterized by high GFA growth that surpasses population demands. S4 follows a rapid economic growth pattern with almost no planning intervention.
3.	 The data set “LUSzoning”.
The dataset includes 16 .asc raster layers providing the simulated land-uses under four defined scenarios for Barcelona, Madrid, Valencia and Zaragoza Functional Urban Areas (FUAs) for 2030. The simulated raster layers were created using CLUMondo simulation framework and have a spatial resolution of 30m. The .asc layers name include the name of the FUA and scenario number. For example, the output from simulating the urban growth for the city of Zaragoza under Scenario 2 is named “Zaragoza_S2.tif”. Furthermore, a .txt file named “Legend.txt” includes the numeric value of the land-use and the category of land-use that represents to interpret the .asc raster layers. The name of the land-use classes is a reclassification of the Urban Atlas 2012 land-use classes within the four Spanish FUAs analyzed.

digital-zoning-plans

display_name: digital-zoning-plans

land-use-change

display_name: land-use-change

land-use-simulations

display_name: land-use-simulations

land-use-transformations

display_name: land-use-transformations

spatial-planning

display_name: spatial-planning

en: LUSzoning: Land-use simulations integrating zoning regulations in Spanish functional urban areas


maintainer: Michael Lehning

en: snow-conservation snowpack snow-farming snow winter-tourism terrestrial-laser-scanning

en: HS Martell autumn 2015

en:
Snow depth/ height of snow heap in autumn 2015 obtained by terrestrial laserscanning
cell size: 10 cm
spatial reference: European Datum 1950 UTM
Nodata: -9999

en: Fluela .smet

en: Meteodata used as input for SNOWAPCK simulations, orginating from Station Windtunnel and DAV.

en: HS Fluela spring 2015

en:
Snow depth/ height of snow heap in spring 2015 obtained by terrrestrial laserscanning
cell size: 10 cm
spatial reference: CH1903

en: Fluela .sno

en: Input snow profile (.sno file) used for SNOWPACK simulations of the Flüela heap

en: Martell .sno

en: Input snow profile (.sno file) used for SNOWPACK simulations of the Martell heap

en: Martell .smet

en: Meteodata used as input for SNOWAPCK simulations, orginating from Station Hintermartell.

en: HS Martell spring 2015

en:
Snow depth/ height of snow heap in spring 2015 obtained by terrestrial laserscanning
cell size: 10 cm
spatial reference: European Datum 1950 UTM
Nodata: -9999

en: HS Fluela autumn 2015

en:
Snow depth/ height of snow heap in autumn 2015 obtained by terrestrial laserscanning
cell size: 10 cm
spatial reference: CH1903
Nodata: -9999

en:
Two data sets obtained for snow farming projects (Fluela, Davos, CH and Martell, IT) in 2015.

The data set contains for each site:

* 10 cm GIS raster of snow depth calculated from terrestrial laserscanning surveys (TLS) in the end of winter season (April/May)

* 10 cm GIS raster of snow depth calculated from TLS in the end of summer season (October)

Input files for SNOWPACK model:

* .sno: snow profile at the end of winter

* .smet: meteorological data measured by weather stations in the area

For more details see Grünewald, T., Lehning, M., and Wolfsperger, F.: Snow farming: Conserving snow over the summer season, The Cryosphere Discuss., https://doi.org/10.5194/tc-2017-93, in review, 2017.

snow

display_name: snow

snow-conservation

display_name: snow-conservation

snow-farming

display_name: snow-farming

snowpack

display_name: snowpack

terrestrial-laser-scanning

display_name: terrestrial-laser-scanning

winter-tourism

display_name: winter-tourism

en: Snowfarming data set Davos and Martell 2015