maintainer: Yohann Chauvier

en: hard-limit conservation large-scale range-maps flags gbif ecoregions landsca species-observations occurrence-records synonyms accepted-name filtering taxonomy species-distribution r-package

en: gbif.range R package version 1.1 (current)

en: gbif.range R package version 0.2 (deprecated)

en: wsl.gbif R package version 0.1 (deprecated)

en: Test version, not to be used.

en: GitHub URL

en: Although species range may be obtained using expert maps or modeling methods, expert data is often species-limited and statistical models need more technical expertise as well as many species observations. When unavailable, such information may be extracted from the Global Biodiversity Information facility (GBIF), the largest public data repository inventorying georeferenced species observations worldwide. However, retrieving GBIF records at large scale may be tedious if users are unaware of specific tools and functions that need to be employed. Here we present *gbif.range*, an R library that contains automated methods to generate species range maps from scratch using in-house ecoregions shapefiles and an easy-to-use GBIF download wrapper. Finally, this library also offers a set of additional very useful parameters and functions for large GBIF datasets (generate doi, extract GBIF taxonomy, records filtering...). [gbif.range R project](https://github.com/8Ginette8/gbif.range)

accepted-name

display_name: accepted-name

conservation

display_name: conservation

ecoregions

display_name: ecoregions

filtering

display_name: filtering

flags

display_name: flags

gbif

display_name: gbif

hard-limit

display_name: hard-limit

landsca

display_name: landsca

large-scale

display_name: large-scale

occurrence-records

display_name: occurrence-records

r-package

display_name: r-package

range-maps

display_name: range-maps

species-distribution

display_name: species-distribution

species-observations

display_name: species-observations

synonyms

display_name: synonyms

taxonomy

display_name: taxonomy

en: gbif.range - An R package to generate species range maps based on ecoregions and a user-friendly GBIF wrapper


maintainer: EnviDat Support

en: european-alps functional-diversity migclim zonation conservation species-distribution-models phylogenetic-diversity beta-diversity taxonomic-diversity global-change land-use-change plants

state: active

type: dataset

en: Data Access

format: GeoTIFF

en: Updated SDM outputs

en: Refined scenarios of plants distribution in the European Alps

en:
This repository contains extensive data for the European Alps:
  - Observations of ~3,500 plant species
  - Climate (1-km), soil (100-m) and land cover predictors (1km); current and future scenarios (28 CMIP6-GCMs, 2 land cover change and 3 dispersal scenarios i.e., unlimited, no and realistic vegetation dispersal)
  - Flora migration rates (categorical) and ecological preferences (continuous indicator values)
  - Regional maps of barriers to migration and water bodies at 100-m resolution
  - Sampling effort, distance to roads and cities predictors at 100-m resolution
  - Present and future abundances over the study region at 1-km resolution (~2,000 species)
  - Present and future multifaceted and uniqueness of the European Alps' Flora at 1-km resolution
  - Present and future conservation recommendations at 1-km resolution (26 current and future strategies)
  - Phylogenetic data and functional traits of ~2,000 plants (raw data and classification trees)
  - All scripts, data and plots used for the analyses, including a singularity container (mini-linux) to run them

beta-diversity

display_name: beta-diversity

european-alps

display_name: european-alps

functional-diversity

display_name: functional-diversity

global-change

display_name: global-change

land-use-change

display_name: land-use-change

migclim

display_name: migclim

phylogenetic-diversity

display_name: phylogenetic-diversity

plants

display_name: plants

species-distribution-models

display_name: species-distribution-models

taxonomic-diversity

display_name: taxonomic-diversity

zonation

display_name: zonation

educ

fr: Éducation, culture et sport

de: Bildung, Kultur und Sport

en: Education, culture and sport

it: Istruzione, cultura e sport

envidat

fr: EnviDat: le portail de données environnementales

de: EnviDat: das Umwelt-Datenportal

en: EnviDat: the Environmental Data Portal

it: EnviDat: il portale dei dati ambientali

fr: EnviDat est le portail de données du WSL; il offre un accès uniformisé et réglementé aux données de surveillance et de recherche en matière d’environnement. Ce portail est capable d’héberger et de publier des ensembles de données précieuses et uniques, qui sont partagées aux quatre coins de la planète. Ce faisant, EnviDat favorise la recherche et contribue à l’avancement des sciences environnementales.

en: EnviDat is the WSL data portal providing unified and managed access to environmental monitoring and research data. The portal has the capability to host and publish data sets. Through the sharing of unique and high-value data sets worldwide, EnviDat fosters research and contributes to advancing environmental science.

de: EnviDat ist das Datenportal der WSL für einen einheitlichen und verwalteten Zugang zu Umweltdaten aus Monitoring- und Forschungsprojekten. Auf dem Portal können Datensätze gehostet und veröffentlicht werden. Durch die weltweite Bereitstellung einzigartiger und wertvoller Datensätze leistet EnviDat einen Beitrag zur Forschung im Bereich der Umweltwissenschaften.

it: EnviDat è il portale dati del WSL che fornisce un accesso unificato e gestito ai dati di monitoraggio ambientale e di ricerca. Il portale ha la capacità di ospitare e pubblicare serie di dati preziosi e unici che vengono condivisi in tutto il mondo. Così facendo, EnviDat promuove la ricerca e contribuisce al progresso della scienza ambientale.

political_level: other

wsl

capacity: public

type: organization

approval_status: approved

key: guid

key: uri

value: Envidat Harvester

en: Present and future multifaceted plant diversity, uniqueness and conservation in the European Alps


en: inter-observer-difference sessile-species detectability lichens biodiversity detection-probability observer-error cryptogams monitoring occupancy-model bayesian-inference swisslichens

media_type: application/zip

en: Repository containing the full data, code, README and workflow files.

format: ZIP

en:
The available lichen data consists of detection/nondetection data (1/0) of 373 tree-inhabiting (*corticolous*) lichen species from 416 plots surveyed 1-2 times. The lichen data were originally collected for the purpose of the Red List of epiphytic lichen species in Switzerland ([Scheidegger *et al.* 2002](https://www.bafu.admin.ch/bafu/de/home/themen/biodiversitaet/publikationen-studien/publikationen/rote-liste-gefaehrdete-arten-baum-erdbewohnende-flechten.html)), but updated to recent nomenclature for the purpose of this study.
This repository contains all the supporting data and R code for the paper:
von Hirschheydt, G., Kéry, M., Ekman, S., Stofer, S., Dietrich, M., Keller, C., Scheidegger, C. (2024) **Occupancy model reveals limited detectability of lichens in a standardised large-scale monitoring**. *Journal of Vegetation Science*.
Results and figures presented in the manuscript should be reproducible (with small differences in the latter digits due to stochasticity of the MCMC sampler) with the provided data and code.
The downloadable `.zip` folder has the following structure:
* `0_data/`
* `1_code/`
* `2_output/`
* `lichen_detectability.Rproj`
* `README.txt`
* `workflow.html`
- `workflow.Rmd`

The main folder and the three subordinate folders each have their own `README*.txt` file. These describe each available file in detail and should be consulted prior to using the data or running any code. The file `lichen_detectability.Rproj` stores the information about the R project. The user can open the project by clicking/double-clicking on this file which will automatically define the repository as working directory for the R session. If the user does not use RStudio/Posit, they may have to set the working directory manually to the stored location in the R files. The files `workflow.*` guide the user through the analysis (`1_code/*.R`) in the correct order so that they can:
- bundle the cleaned data into a data list readable for JAGS
- fit the multi-species occupancy model to the data and store the output
- assess the goodness-of-fit of the model to the data
- conduct a prior sensitivity analysis with 2 additional sets of priors
- extract the summary statistics reported in the manuscript and supplementary materials
- generate the figures shown in the manuscript and supplementary materials

bayesian-inference

display_name: bayesian-inference

biodiversity

display_name: biodiversity

cryptogams

display_name: cryptogams

detectability

display_name: detectability

detection-probability

display_name: detection-probability

inter-observer-difference

display_name: inter-observer-difference

lichens

display_name: lichens

monitoring

display_name: monitoring

observer-error

display_name: observer-error

occupancy-model

display_name: occupancy-model

sessile-species

display_name: sessile-species

swisslichens

display_name: swisslichens

en: Repeated detection-nondetection data of corticolous lichens from a standardised monitoring across Switzerland


causal-effect-of-mfoz

title_for_slug: causal-effect-of-mfoz

en: built-up-land-change causality the-major-function-oriented-zone fujian-province selection-bias propensity-score-matching

en: Data and Stata code for repeating the results

en:
Data.xlsx is the variables of 952 townships
Table 1-2 NN(1-1)_noreplace_caliper(0.01).do is STATA code to produce the results in Table 1 and 2
Figure 5 Distribution of the propensity scores.R is R code rto produce Figure 5
Figure 5 Standard mean difference.R is R code rto produce Figure 5

en:
Title: Closer to causality: How effective is spatial planning in governing built-up land expansion in Fujian Province, China?

Research objective: The Major Function Oriented Zone (MFOZ), the first strategic spatial plan in China, is developed to achieve a coordinated regional development, through spatial regulation and zoning of development. The MFOZ he MFOZ divided land into four major function-oriented zones: The development-optimized zone, the development-prioritised zone, the development-restricted zone, and the development-prohibited zone. We used propensity score marching to evaluate the effect of the MFOZ on built-up land expansion in Fujian Province over three time intervals (2013–2015, 2013–2018 and 2013–2020).

Data: Data.xlsx contains the variables of 954 towns in Fujian Province.
Town_ID is the town unique ID;
County_ID is the county unique ID;
City_ID is the city unique ID;
MFOZ is the the development-prioritised zone and the development-restricted zone (The development-optimized zone and the development-prohibited zone are excluded);
Builtup_13_15 is the built-up land expansion from 2013 to 2015;
Builtup_13_18 is the built-up land expansion from 2013 to 2018;
Builtup_13_20 is the built-up land expansion from 2013 to 2020;
Dis2water is the Euclidean distance from the town to the nearest waterbody;
Slope is the the average slope within the town;
GDP is the average GDP in 2010 within the town;
Pop is the average population in 2010 within the town;
Road is the average population in 2010 within the town;
Dis2city is the Euclidean distance from the town to the nearest prefectural city centre;
Nei_Arable, Nei_Forest, and Nei_Built.up are the area of arable land, forest land, and built-up land neighbouring town i in 2010.

Method: we used the propensity score matching to compare the changes in the amount of built-up land in the towns of the development-prioritised zone with the matched towns of the development-restricted zone. Additionally, we used three evaluation intervals (2013–2015, 2013–2018 and 2013–2020) to evaluate temporal variation in the causal effect of the MFOZ on built-up land expansion.

built-up-land-change

display_name: built-up-land-change

causality

display_name: causality

fujian-province

display_name: fujian-province

propensity-score-matching

display_name: propensity-score-matching

selection-bias

display_name: selection-bias

the-major-function-oriented-zone

display_name: the-major-function-oriented-zone

en: Causal effect of MFOZ


backward-trajectories

maintainer: Michael Sprenger

title_for_slug: backward-trajectories

en: complex-topopgraphy cosmo lagrangian-analysis raclets alps atmospheric-dynamics backward-trajectories

en: Backward Trajectories

en: The trajectories are started from surface up to 5000 m altitude, with an equidistant vertical spacing of 200 m, and calculated 120 h backward in time. As long as the trajectories stay in the COSMO1 domain (essentially the Alpine region), as run operationally at MeteoSwiss, the COSMO analysis wind fields are used. When the trajectories leave this domain, they are extended further backward in time based on ECMWF analysis data. The following fields are traced along the trajectories: longitude, latitude, geopotential height, pressure, specific humidity, temperature and surface topography. Furthermore, it is indicated for each trajectory time instance on which model data, COSMO or ECMWF, the trajectory time is based. Finally, the trajectories are started every hour in the specified time period, and positions and fields are available every 10 min. Each day is saved as a separate file. The last 8 digits of the file names indicate the starting date of the trajectories.

mimetype: application/zip

en: Backward trajectories were calculated from two positions: Davos Wolfgang (LON: 9.85361, LAT: 46.83551) and Weissfluhjoch (LON: 9.80646 LAT: 46.83304) for the time period February 2 until March 27 2019 using COSMO or ECMWF, respectively.

alps

display_name: alps

atmospheric-dynamics

display_name: atmospheric-dynamics

backward-trajectories

display_name: backward-trajectories

complex-topopgraphy

display_name: complex-topopgraphy

cosmo

display_name: cosmo

lagrangian-analysis

display_name: lagrangian-analysis

raclets

display_name: raclets

en: Backward  Trajectories


maintainer: Esther Frei

en: long-term-experiment literature-review treeline-ecosystems mountain-ecology stillberg

en: The Stillberg bibliography (Stillberg_bibliography_v1.zip) comprises a comprehensive list of 276 scientific publications, 91 of them published in peer-reviewed ISI journals (Stillberg_bibliography_data_v1.csv) along with the metadata file (Stillberg_bibliography_metadata_v1.csv). Currently the bibliography comprises literature about the main afforestation experiment, the FACE × warming experiment, the nutrient addition experiment, and the G-TREE experiment, as well as further publications related to the Stillberg research site that have been published until August 2023. The bibliography can be filtered for different categories, e.g., experiment, peer-review, source repository or database, and source title.

format: SERVICE

en:
 Background information

The Stillberg ecological treeline research site in the Swiss Alps was established in 1975, with the aim to develop ecologically, technically, and economically sustainable reforestation techniques at the treeline to reduce the risk of snow avalanches. In the course of time, additional research aspects gained importance, such as the ecology of the treeline ecotone under global change. Over almost fifty years, research at the Stillberg site combined long-term monitoring of the large-scale high-elevation afforestation with experimental manipulations simulating global change impacts. Besides providing a scientific basis and practical guidelines for high-elevation afforestation, this research has contributed to a comprehensive understanding of ecological processes in the treeline ecotone across different compartments and scales, from individual trees, non-tree vegetation and soils to whole ecosystems, in the context of global change resulting in more than 150 publications.

 Dataset generation

We compiled a comprehensive list of scientific publications covering research at the Stillberg research site by conducting searches in the literature databases Web of Science and Google Scholar, as well as in the Digital Object Repository of the Swiss Federal Institute for Forest, Snow and Landscape Research WSL (DORA). We compiled all publications about the afforestation experiment, the FACE × warming experiment, the nutrient addition experiment, the G-TREE experiment, as well as other studies related to the Stillberg research site.

 Data description

The Stillberg bibliography (Stillberg_bibliography_data_v1.csv) comprises a comprehensive list of 276 scientific publications, 91 of them published in peer-reviewed ISI journals. Currently the bibliography comprises literature about the main afforestation experiment, the FACE × warming experiment, the nutrient addition experiment, and the G-TREE experiment, as well as further publications related to the Stillberg research site that have been published until August 2023. The bibliography can be filtered for different categories, e.g., experiment, peer-review, source repository or database, and source title. The bibliography is described in a metadata file (Stillberg_bibliography_metadata_v1.csv). The bibliography along with the metadata file are provided in a ZIP-folder (Stillberg_bibliography_v1.zip).

literature-review

display_name: literature-review

long-term-experiment

display_name: long-term-experiment

mountain-ecology

display_name: mountain-ecology

stillberg

display_name: stillberg

treeline-ecosystems

display_name: treeline-ecosystems

educ

en: Education, culture and sport

envidat

en: EnviDat: the Environmental Data Portal

en: Bibliography of the long-term treeline research site Stillberg, Switzerland


maintainer: Athanasios Nenes

en: wolfgangpass supersaturation cloud-droplets sigma-w raclets cloud-modeling

en: Predicted cloud droplet numbers (kappa=0.25) Weissfluhjoch

media_type: text/csv

en:
In-situ measurements of aerosol size distributions along with updraft velocities collected from a wind Doppler Lidar are coupled with a state-of-the art droplet activation parameterization (Morales and Nenes, 2014) to determine the cloud droplet number and maximum in-cloud supersaturation that can potentially form over the Davos Wolfgang station between February 24 and March 8 2019. For this, as a sensitivity test, a hygroscopicity parameter of 0.25 is assumed, which is the upper limit determined from the analysis carried out for Weissfluhjoch (https://www.envidat.ch/dataset/ccn-hygroscopicity-predicted-cloud-droplet-numbers-weissfluhjoch). The data file contains:

* Date and time (DD/MM/YYYY hh:mm:ss) in UTC
* The wind-lidar derived standard deviation of updraft velocities (sigma-w) in m/s, used to apply the parameterization employing a single characteristic velocity (Morales and Nenes, 2010)
* The maximum predicted cloud parcel supersaturation (Smax%)
* The total in-situ aerosol number concentration (Naer) in cmˉ³, measured by the SMPS
* The predicted cloud droplet numbers (Nd) in cmˉ³
* The activation fraction (%)

format: CSV

mimetype: text/csv

en: Predicted cloud droplet numbers (kappa=0.1) Davos Wolfgang

en:
In-situ measurements of aerosol size distributions along with updraft velocities collected from a wind Doppler Lidar are coupled with a state-of-the art droplet activation parameterization (Morales and Nenes, 2014) to determine the cloud droplet number and maximum in-cloud supersaturation that can potentially form over the Davos Wolfgang station between February 24 and March 8 2019. For this, as a sensitivity test, a hygroscopicity parameter of 0.1 is assumed, which is the lower limit determined from the analysis carried out for Weissfluhjoch (https://www.envidat.ch/dataset/ccn-hygroscopicity-predicted-cloud-droplet-numbers-weissfluhjoch). The data file contains:

* Date and time (DD/MM/YYYY hh:mm:ss) in UTC
* The wind-lidar derived standard deviation of updraft velocities (sigma-w) in m/s, used to apply the parameterization employing a single characteristic velocity (Morales and Nenes, 2010)
* The maximum predicted cloud parcel supersaturation (Smax%)
* The total in-situ aerosol number concentration (Naer) in cmˉ³, measured by the SMPS
* The predicted cloud droplet numbers (Nd) in cmˉ³
* The activation fraction (%)

en: Cloud droplet properties were predicted between February 24 and March 8 2019 for the measurement site Davos Wolfgang (1630 m a.s.l., LON: 9.853594, LAT: 46.835577). Droplet calculations are carried out with the physically based aerosol activation parameterization of Morales and Nenes (2014), employing the “characteristic velocity” approach of Morales and Nenes (2010). Aerosol size distribution observations required to predict the cloud droplet numbers and maximum in-cloud supersaturation are obtained from a Scanning Mobility Particle Size Spectrometer (SMPS) instrument deployed at Davos Wolfgang (https://www.envidat.ch/dataset/aerosol-data-davos-wolfgang). The required vertical velocity measurements are derived from the wind Doppler Lidar (https://www.envidat.ch/dataset/lidar-wind-profiler-data) deployed at the same station and are extracted from the first bin of the instrument, being 200 m above ground level. The hygroscopic properties of the particles measured at Davos Wolfgang could not be estimated, owing to a lack of concurrent CCN measurements. As a sensitivity test, droplet calculations are performed assuming two different values of the aerosol hygroscopicity parameter, 0.1 and 0.25, based on the analysis carried out for Weissfluhjoch. Additional information can be found in Section 2.3 [here](https://acp.copernicus.org/preprints/acp-2020-1036/).

cloud-droplets

display_name: cloud-droplets

cloud-modeling

display_name: cloud-modeling

sigma-w

display_name: sigma-w

supersaturation

display_name: supersaturation

wolfgangpass

display_name: wolfgangpass

en: EnviDat is the WSL data portal providing unified and managed access to environmental monitoring and research data. The portal has the capability to host and publish data sets. Through the sharing of unique and high-value data sets worldwide, EnviDat fosters research and contributes to advancing environmental science.

en: Predicted cloud droplet numbers Davos Wolfgang


maintainer: Anita C. Risch

en: nutrient-network soil grassland global mineralization worldwide soil-biology soil-chemistry

en:
site.code: Site code

Continent: Continent

sampleID: Sample ID

nn.latitude: Lattitude

nn.longitude: Longitude

bioclim.T.ann: Mean annual temperature (°C)

bioclim.T.var: Temperature variability (°C)

bioclim.T.q.wet: Temperature of the wettest quarter (°C)

bioclim.T.q.dry: Temperature of the driest quarter (°C)

bioclim.T.q.warm: Temperature of the warmest quarter (°C)

bioclim.P.ann: Mean annual preciptiation (mm)

bioclim.P.q.wet: Precipitation of the wettest quarter (mm)

ch.pH: pH

ch.WSK.vol: Water holding capacity (vol%)

ch.bulkdensity.gcm3: Bulk density (g cm-3)

ch.porespace: Pore space (vol%)

ch.sand.perc: Soil texture: sand (%)

ch.silt.perc: Soil texture: silt (%)

ch.clay.perc: Soil texture: clay (%)

ch.NO3start.mgg: NO3 at the beginning of the incubation (mgkg)

ch.Ntot.perc: Total soil N (%)

ch.Ctot.perc: Total soil C (%)

ch.Corg.perc: Soil organic C (%)

ch.netNminfield.mgkgd: Realised soil net N mineralisation (mg kg-1 d-1)

ch.netNminlab.mgkgd: Potential soil ne N mineralisation (mg kg-1 d-1)

archaea.cpg.log: Ammonia oxidising archaea (archaeal amoA gene copies g-1 soil)

bacteria.cpg.log: Ammonia oxidising bacteria (bacterial amoA gene copies g-1 soil)

micr.biom: Microbial biomass (ug g-1 soil)

ch.CtoN: C to N ratio

ch.netNnitriffield.mgkgd.sqrt: Realised soil net nitrification, transformed (mg kg-1 d-1)

ch.netNammoniffield.mgkgd.sqrt: Realised soil net ammonification, transformed (mg kg-1 d-1)

ch.netNnitriflab.mgkgd.sqrt: Potential soil net nitrification, transformed (mg kg-1 d-1)

ch.netNammoniflab.mgkgd.sqrt: Potential soil net ammonifiation, transformed (mg kg-1 d-1)

format: XLS

en:
This dataset contains all data on which the following publication below is based.

Paper Citation:

Risch, A. C.; Zimmermann, S.; Ochoa-Hueso, R.; Schütz, M.; Frey, B.; Firn, J. L.; Fay, P. A.; Hagedorn, F.; Borer, E. T.; Seabloom, E. W.; et al. Soil net nitrogen mineralisation across global grasslands. Nat. Commun. 2019, 10 (1), 4981 (10 pp.). doi.org/10.1038/s41467-019-12948-2

Please cite this paper together with the citation for the datafile.

We conducted coordinated measurements of realised and potential soil net Nmin, and assessed water holding capacity, bulk density, C and N content, texture, pH, pore space, microbial biomass, and archaeal (AOA) and bacterial (AOB) ammonia oxidiser abundance using identical materials and methods across 30 grasslands on six continents. The sites covered a globally relevant range of climatic and edaphic conditions. Climate data was obtained from worldclim - Global climate data https://www.worldclim.org/

global

display_name: global

grassland

display_name: grassland

mineralization

display_name: mineralization

nutrient-network

display_name: nutrient-network

soil

display_name: soil

soil-biology

display_name: soil-biology

soil-chemistry

display_name: soil-chemistry

worldwide

display_name: worldwide


en: spatial-planning land-use-change land-use-transformations strategic-spatial-planning urban-transformations europe large-scale-urban-development-projects strategic-projects

en: QCA data set large-scale urban development projects

media_type: text/plain

en: Please refer to the sections: 3. The data set lsUDPs: 3.1 Selection of cases and data collection; 3.2 Data management and operationalization in the Metadata for further details on this resource.

format: TXT

mimetype: text/plain

en: QCA R script large-scale urban development projects Europe

en:
For the set up of this R script, we used the “QCA” (Duşa, 2019) and “SetMethods” (Oana and Schneider, 2018) R packages.

The following further resources were also used:
Dusa, A. (2019). QCA with R: A comprehensive resource. Springer: Berlin.
Schneider, C. Q., & Wagemann, C. (2012). Set-theoretic methods for the social sciences. A guide to qualitative comparative analysis. Cambridge: Cambridge University Press.
Thomann, E., Oana, I.-E., & Wittwer, S. (2018). Performing fuzzy- and crisp set QCA with R. A user-oriented beginner's guide. Retrieved from http://www.evathomann.com/links/qca-r-manual

en:
Table of Content: 1. General context of the data set "lsUDPs" ; 2. Background and aims of the study using the data set lsUDPs; 3. The data set lsUDPs: 3.1 Selection of cases and data collection; 3.2 Data management and operationalisation

1. General context of the data set "lsUDPs"
The data set "lsUDPs" has been generated as part of the CONCUR research project (https://www.wsl.ch/en/projects/concur.html) led by Dr. Anna M. Hersperger and funded by the Swiss National Science Foundation (ERC TBS Consolidator Grant (ID: BSCGIO 157789) for the period 2016-2020. The CONCUR research project is interdisciplinary and aims to develop a scientific basis for adequately integrating spatial policies (in this case, strategic spatial plans) into quantitative land-change modelling approaches at the urban regional level. The first stage (2016-2017) of the CONCUR project focussed on 21 urban regions in Western Europe. The urban regions were selected through a multi-stage strategy for empirical research (see Hersperger, A. M., Grădinaru, S., Oliveira, E., Pagliarin, S., & Palka, G. (2019). Understanding strategic spatial planning to effectively guide development of urban regions. Cities, 94, 96–105. https://doi.org/10.1016/j.cities.2019.05.032 ).

2. Background and aims of the study using the data set lsUDPs
As part of the CONCUR project, a specific task was to examine the relationship between strategic spatial plans and the formulation and implementation (i.e. urban land change) of large-scale urban development projects in Western Europe.
Strategic urban projects are typically large-scale, prominent urban transformations implemented locally with the aim to stimulate urban growth, for instance in the form of urban renewals of deprived neighborhoods, waterfront renewals and transport infrastructures. While strategic urban projects are referred to in the literature with multiple terms, in the CONCOR project we call them large-scale urban development projects (lsUDPs).
Previous studies acknowledged both local and supra-local (or structural) factors impacting the context-specific implementation of lsUDPs. Local governance factors, such as institutional capacity, coordination among public and private actors and political leadership, intertwine with supra-local conditions, such as state re-scaling processes and devolution of state competencies in spatial planning, de-industrialisation and increasing social inequality. Hence, in implementing lsUDPs, multi-scalar factors act in combination.
Because the formulation and implementation of lsUDPs require multi-scalar coordination among coalitions of public and private actors over an extended period of time, they are generally linked to strategic spatial plans (SSPs). Strategic spatial plans convey collective visions and horizons of action negotiated among public and private actors at the local and/or regional level to steer future urban development, and can contain legally binding dispositions, but also indicative guidelines.
The key question remains as to what extent large-scale urban development projects and strategic spatial plans can be regarded as aligned. By alignment, or “concordance”, we mean that strategic projects are formulated and implemented as part of the strategic planning process (“high concordance”), or that the strategic role of projects is reconfirmed in (subsequent) strategic plans (“moderate concordance”). Lack of concordance is found when lsUDPs have been limitedly (or not at all) acknowledged in strategic spatial plans.
We assume that certain local and supra-local factors, characterising the development of the projects, foster (but not strictly “cause”) the degree of alignment between lsUPDs and SSPs. In this study, we empirically examine how, and to what extent, the concordance between 38 European large-scale urban development projects and strategic plans (outcome: CONCOR) has been enabled by five multi-scalar factors (or conditions): (i) the role of the national state (STATE), (ii) the role of (inter)national private actors (PRIVATE), (iii) the occurrence of supra-regional external events (EVENTS), (iv) the degree of transport connectivity (TRANSP), and (v) local resistance from civil society (RESIST). We adopted a (multi-data) case-based qualitative strategy for empirical research and applied the formalised procedure of within- and cross-case comparison offered by fuzzy-set Qualitative Comparative Analysis appropriate for the goal of this study. Based on set theory, QCA formally integrates contextual sensitivity to case specificities (within-case knowledge) with systematic comparative analysis (across-case knowledge).
The research question the data set has been created to reply to is the following: which conditions, and combinations of conditions, enable the concordance between large-scale urban development projects and strategic spatial plans?
The conditions (“independent variables”) considered are. STATE: the set of large-scale urban projects characterized by a high degree of state intervention and support in their formulation and implementation, PRIVATE: the set of large-scale urban projects characterized by a high degree of involvement of (inter)national private actors in their formulation and implementation, EVENTS: the set of large-scale strategic projects whose formulation and implementation have been strongly affected by unforeseen international events and/or global trends, TRANSP: the set of large-scale strategic projects with a high degree of road and/or transit connectivity, and RESIST: set of large-scale strategic projects whose realization has been characterized by resistances that have substantially delayed or modified the project implementation.
The outcome (“dependent variable”) under analysis is CONCOR: the set of large-scale urban projects having a high degree of concordance/alignment/integration with strategic spatial plans

3. The data set lsUDPs

   3.1 Selection of cases and data collection
To generate the current data set on large-scale urban development projects in European urban regions (data set "lsUDPs"), we identified 35 large-scale urban development projects in a sample of the 21 Western urban regions considered in the CONCUR project (see supra, Hersperger et al. 2019): Amsterdam, Barcelona, Copenhagen, Hamburg, Lyon, Manchester, Milan, Stockholm, Stuttgart. The criteria we followed to identify the 35 large-scale urban development projects are: geographical location, size (large-scale), site (located either in the city core or in the larger urban region) and urban function (e.g. housing, transportation infrastructures, service and knowledge economic functions). Employing these criteria facilitated the selection of diverse large-scale urban development projects while still ensuring sufficient comparability.
In 2016, we performed 47 in-depth interviews with experts in urban and regional planning and large-scale strategic projects and infrastructure (i.e. academics and practitioners) about the formulation, implementation and development (1990s–2010s) of each project in each of the 9 selected urban regions. On average, each interviewee answered questions on 3.1 large-scale urban development projects. Three cases were subdivided into two cases because a clear differentiation between specific implementation stages was identified by the interviewees (expansion of the Barcelona airport, cases “bcn_airport80-90” and “bcn_airport00-16”; realisation of Lyon Part-Dieu, cases “lyo_partdieu70-90” and “lyo_partdieu00-16”; MediaCityUK, cases “man_salfordquays80-00” and “man_mediacityuk00-16”). Therefore, from the initial 35 cases, the final number of analysed cases in the lsUDPs dataset is 38.

    3.2 The data set lsUDPs: Data management and operationalisation
Interviews were fully transcribed and analysed through MAXQDA (version 12.3, VERBI GmbH, Berlin, Germany), and intercoder agreement was evaluated on a sample of nine interviews. We also compiled “synthetic case descriptions” (SCD) for each case (totalling more than 160 SCDs) to spot potential inconsistencies among interviewees’ accounts and to facilitate completion of the “calibration table” for each case (see below). An online expert survey distributed to the interviewees (response rate 78%) helped systematise the information collected during the interviews. We also consulted both academic and gray literature on the case studies to check for possible ambiguity and inconsistencies in the interview data, and to solve discrepancies between our assigned set membership scores and questionnaire values. Site visits were also carried out to retrieve additional information on the selected cases.
For each case (i.e. each of the 38 selected large-scale urban development projects), we operationalised each condition (i.e. STATE, PRIVATE, EVENTS, TRANSP, RESIST) and the outcome (CONCOR) in terms of sets, for subsequent application of Qualitative Comparative Analysis. This process is called “calibration”; we used a number of indicators for each condition to qualitatively assess each large-scale project across the conditions. The case-based qualitative assessment was then transformed into fuzzy-set membership values. Fuzzy-set membership values range from 0 to 1, and should be conceived as “fundamentally interpretative tools” that “operationalize theoretical concepts in a way that enhances the dialogue between ideas and evidence” (Ragin 2000:162, in “Fuzzy-set Social Science”. Chicago: University Press). We employed a four-value fuzzy-set scale (0, 0.33, 0.67, 1) to “quantify” into set membership scores the individual histories of cases retrieved from interview data. Only the condition TRANSP was calibrated as a crisp-set (0, 1).
The translation of qualitative case-based information into numerical fuzzy-set membership values was iteratively performed by populating a calibration table following standard practices recently emerged in QCA when dealing with qualitative (interview) data.

europe

display_name: europe

land-use-change

display_name: land-use-change

land-use-transformations

display_name: land-use-transformations

large-scale-urban-development-projects

display_name: large-scale-urban-development-projects

spatial-planning

display_name: spatial-planning

strategic-projects

display_name: strategic-projects

strategic-spatial-planning

display_name: strategic-spatial-planning

urban-transformations

display_name: urban-transformations

en: lsUDPS Large-scale urban development projects in European urban regions


high-resolution-climate-data-for-europe

title_for_slug: high-resolution-climate-data-for-europe

en: eur11 europe air-temperature daily-climate minimum-temperature cordex maximum-temperature precipitation

en: CHELSA EUR11 Technical Documentation

media_type: application/pdf

en: As the CHELSA project is continuously expanded, certain new parameters, time scales etc. are not included anymore in the original publication (Karger et al. 2017). We are therefore provide this document as a guideline of the current climatic parameters available and will expand it as new parameters or time periods will become available. Please refer to the version history for changes made in the document.

format: PDF

en: Data Access

en:
High-resolution information on climatic conditions is essential to many applications in environmental and ecological sciences. Here we present downscaled climate data for the CORDEX EUR11 domain at a high resolution of 30 arc sec. The temperature algorithm is based on statistical downscaling of atmospheric temperature lapse rates. The precipitation algorithm incorporates orographic predictors including wind fields, valley exposition, and boundary layer height. The resulting data consist of a daily temperature and precipitation timeseries.

The data is distributed under a: Creative Commons: Attribution 4.0 International (CC BY 4.0) license.

air-temperature

display_name: air-temperature

cordex

display_name: cordex

daily-climate

display_name: daily-climate

maximum-temperature

display_name: maximum-temperature

minimum-temperature

display_name: minimum-temperature

precipitation

display_name: precipitation

educ

en: Education, culture and sport

envidat

en: EnviDat: the Environmental Data Portal

en: High resolution climate data for Europe


en: acoustic-emissions snow fiber-bundle-model sintering viscous-deformation snow-failure snow-avalanche snow-mechanics

en: fiber-bundle-model-for-snow-failure

en: Data underlying Figures in publication "Modelling Snow Failure Behavior and Concurrent Acoustic Emissions Signatures with a Fiber Bundle Model" submitted to "Geophysical Research Letters"

en:
This dataset contains modeled and experimental results for laboratory snow failure experiments and the concurrent acoustic emissions signatures for different loading rates. For modelling the snow failure we used a fiber bundle model that includes sintering and viscous deformation.

The data underlay the figures in the publication "Modelling Snow Failure Behavior and Concurrent Acoustic Emissions Signatures with a Fiber Bundle Model" submitted for publication to "Geophysical Research Letters".

acoustic-emissions

display_name: acoustic-emissions

fiber-bundle-model

display_name: fiber-bundle-model

sintering

display_name: sintering

snow

display_name: snow

snow-avalanche

display_name: snow-avalanche

snow-failure

display_name: snow-failure

snow-mechanics

display_name: snow-mechanics

viscous-deformation

display_name: viscous-deformation

envidat

en: EnviDat is the WSL data portal providing unified and managed access to environmental monitoring and research data. The portal has the capability to host and publish data sets. Through the sharing of unique and high-value data sets worldwide, EnviDat fosters research and contributes to advancing environmental science.

en: Fiber Bundle Model for snow failure and concurrent Acoustic Emissions


en: growth-reaction-to-drought-stress forests fagus-sylvatica european-summer-drought drought-2018 leaf-senescence

en: Drought and Beech: 1000 beeches data set

en: The data set comprises data from the Drought and Beech 1000 beeches project. The actual data file with monitoring data of 963 beech trees in northern Switzerland over four years (Drought_and_Beech_1000beeches_Data_v1.0.csv), a metadata file with column descriptions (Drought_and_Beech_1000beeches_Legend_v1.0.csv), and a file defining the different survey periods (Drought_and_Beech_1000beeches_Survey_periods_v1.0.csv).

en: This study investigated multi-year drought impacts on beech forests through a unique large-scale monitoring of 963 individual beech trees, which showed either premature leaf discoloration during the drought in summer 2018 or no visible damage. We conducted the study in two highly drought-affected regions in northern Switzerland and one less drought-affected region located further south. We quantified the development of crown dieback and tree mortality as well as secondary drought damage, i.e. the presence of bleeding cankers and bark beetle infestations, in these trees for three consecutive years. We also determined the impact of several potential climate- and stand-related (predisposing) factors on mortality and drought legacy processes.

european-summer-drought

display_name: european-summer-drought

fagus-sylvatica

display_name: fagus-sylvatica

forests

display_name: forests

growth-reaction-to-drought-stress

display_name: growth-reaction-to-drought-stress

leaf-senescence

display_name: leaf-senescence

en: Data on multi-year drought impacts on European beech in northern Switzerland


swiss-fluxnet-site-davos

title_for_slug: swiss-fluxnet-site-davos

en: eddy-covariance swiss-forest-lab greenhouse-gas flux swiss-fluxnet meteorological-measurements icos forest

en: Processed data

en: Processed flux and meteo data are available from the European Fluxes Database Cluster.

en: Fluxnet2015 dataset

en: Processed flux and meteo data are also part of the Fluxnet2015 dataset.

en: Site description

en: Data access

en: The flux and meteo data of the Swiss FluxNet site Davos are openly available. Near real-time flux and meteo data are uploaded daily to the ICOS Carbon Portal.

en:
The Swiss FluxNet Site Davos is a managed subalpine evergreen forest, located on the Seehorn mountain near Davos in the Swiss Alps. The site is dominated by Norway spruce. The tower is owned by the Federal Office for the Environment (FOEN).
Ecosystem flux measurements of CO2, H2O (since 1997) as well as CH4 and N2O (since 2016) are performed with the eddy covariance method.
In addition to Swiss FluxNet, the site is part of the National Air Pollution Monitoring Network (NABEL), the Long term Forest Ecosystem Research (LWF), the biological drought and growth indicator network (TreeNet) and of ICOS Switzerland (Integrated Carbon Observation System). Since November 2019, the site is an ICOS Class 1 Ecosystem station.

Measurements

- Ecosystem flux measurements of CO2, H2O vapour (since 1997) as well a CH4 and N2O (since 2016) are performed with the eddy-covariance method. This method is based on measurements of trace gas mixing ratios, using infrared gas analyzers (for CO2, H2O vapor) and laser spectrometers (for CH4 and N2O), combined with wind speed and wind direction measurements, using 3D sonic anemometers. To resolve the short-term turbulent fluctuations in the atmosphere, very fast measurements are needed: we measure at 10-20 Hz, i.e., 10-20 times per second. To assess the energy budget of each ecosystem, also radiation sensors and soil climate profiles are installed at the site.
- Sub-canopy eddy fluxes (CO2, H2O, since 2023 also CH4).
- Continuous profile concentration and forest floor flux measurement of CO2, H2O, CH4, N2O.
- Auxiliary micrometeorology and soil climate measurements.

Data availability

Near real-time flux and meteo data uploaded daily to the ICOS Carbon Portal. Processed flux and meteo data are also available from the European Fluxes Database Cluster and part of Fluxnet2015 dataset.

Data policy

ICOS data license:
[https://www.icos-cp.eu/data-services/about-data-portal/data-license](https://www.icos-cp.eu/data-services/about-data-portal/data-license)

Detailed site info:
[https://www.swissfluxnet.ethz.ch/index.php/sites/ch-dav-davos/site-info-ch-dav/](https://www.swissfluxnet.ethz.ch/index.php/sites/ch-dav-davos/site-info-ch-dav/)

eddy-covariance

display_name: eddy-covariance

flux

display_name: flux

forest

display_name: forest

greenhouse-gas

display_name: greenhouse-gas

icos

display_name: icos

meteorological-measurements

display_name: meteorological-measurements

swiss-fluxnet

display_name: swiss-fluxnet

swiss-forest-lab

display_name: swiss-forest-lab

en: Swiss FluxNet Site Davos


maintainer: Dirk Nikolaus Karger

en: species-distribution-models iucn generalized-additive-models amphibians sdm mammals interannual-climate-variability range-maps climate-variablity random-forests niche-models reptiles chelsa generalized-linear-models

en: Readme file containing the file naming convention, file dimensions, and variable names

en: File containing the climate predictors as SAGA grid files (.sgrd)

en: Reptiles

en: Species distribution models based on different climate predictor groups

en: Amphibians

en: File containing the record dimension, genus, and species of the respective netcdf files

en: Mammals

en: We modelled the global distribution of 730 amphibian, 1276 reptile, and 1961 mammal species globally as a function of current climate at a 0.5° spatial resolution using four different predictor groups composed of different combinations of input variables: mean climatic conditions, spatial climatic variability, and temporal (interannual) climatic variability.

amphibians

display_name: amphibians

chelsa

display_name: chelsa

climate-variablity

display_name: climate-variablity

generalized-additive-models

display_name: generalized-additive-models

generalized-linear-models

display_name: generalized-linear-models

interannual-climate-variability

display_name: interannual-climate-variability

iucn

display_name: iucn

mammals

display_name: mammals

niche-models

display_name: niche-models

random-forests

display_name: random-forests

range-maps

display_name: range-maps

reptiles

display_name: reptiles

sdm

display_name: sdm

species-distribution-models

display_name: species-distribution-models

educ

en: Education, culture and sport

envidat

en: EnviDat: the Environmental Data Portal

en: EnviDat is the WSL data portal providing unified and managed access to environmental monitoring and research data. The portal has the capability to host and publish data sets. Through the sharing of unique and high-value data sets worldwide, EnviDat fosters research and contributes to advancing environmental science.

en: Global species distributions for mammals, reptiles, and amphibians


maintainer: Jana von Freyberg

en: precipitation catchment hydrology stable-water-isotope alptal stream-water

en:
Version 2 of the dataset contains additional information on:
- line-conditioned (lc) excess
- precipitation sample volumes
- isotope data quality

en: Data Set Documentation: alptal_Isotopes_daily_2015-2019_v2

en: Documentations contains meta data of the data columns and information about the study catchments.

en: Single comma-separated txt-file with daily data of stable water isotopes measured in streamwater and precipitation in the Swiss Alptal catchment and two of its tributaries (Erlenbach and Vogelbach), as well as daily data of key hydrometeorological variables from the Alptal valley

en: This dataset contains four years of daily measurements of the natural isotopic composition (2H, 18O) of precipitation and stream water at the Alp catchment (area 47 km2) in Central Switzerland and two of its tributaries (0.73 km2 and 1.55 km2). In addition, the dataset contains daily measurements of key hydrometeorological variables.

alptal

display_name: alptal

catchment

display_name: catchment

hydrology

display_name: hydrology

precipitation

display_name: precipitation

stable-water-isotope

display_name: stable-water-isotope

stream-water

display_name: stream-water

en: Four years of daily stable water isotope data in stream water and precipitation from three Swiss catchments


en: sea-salt-aerosol snow arctic antarctica ice

en: Fig 1F_1

en: Individual data to mean shown in Figure 1F

en: Fig 1F

en: Data to Figure 1F

en: Fig 1B_2

en: Individual data to mean shown in Figure 1B

en: Fig 1F_2

en: Fig 1C

en: Data to Figure 1C

en: Fig 1E

en: Data to Figure 1E

en: Fib 1B_3

en: This table lists all data files in chronological order. "SampleTemp" refers to the reading of the PT-1000 at the side of the sample holder. "cellPressure" is the water vapour pressure during the measurements. From this, in presence of ice, the temperature of the ice sample can be calculated which is given in the column "iceTemp". "dT" is the difference, or temperature offset, between sensor's reading and the ice samples temperature. The offset is due to the sensor being mounted away from the cooled spot of the sample were it experiences heating from warmer parts of the sample holder. The off-set is used to calibrate the temperature sensor and the resulting temperature during the experiments is given in the column "temperature". "relHum" is the relative humidity above (supercooled) liquid and "concBrine" is the concentration of a NaCl solution in mol/l in equilibrium with ice at the experimental temperature. "Area" is the integrated PE signal.

en: Read Me

en: Description of the data tables

mimetype: application/pdf

en: Fig 1D

en: Data to Figure 1D

en: Fig 1A

en: Data Figure 1A

en: Code

en: Matlab code to read and plot the data

en: Table 2

en: This table lists all NEXAFS data files in chronological order with PE carbon to oxygen signal intensity ratios -from which the atomic ratio were calculated in the paper - before ("preCtoO") and after ("postCtoO") each NEXAFS measurement. Also listed is the change in Na ("lossSodi"), O ("lossOxy"), Cl ("lossChlor"), and C ("lossCarb") signal intensity after the NEXAFS measurement compared to prior to the NEXAFS measurement.

en: Fig 1B_1

en: Laboratory experiments are presented on the phase change at the surface of sodium chloride – water mixtures at temperatures between 259 K and 240 K. High selectivity to the upper few nanometres of the frozen solution – air interface is achieved by using electron yield near-edge X-ray absorption fine structure (NEXAFS) spectroscopy. We present the NEXAFS spectrum of the hydrohalite.

en: Data set on interfacial supercooling and the precipitation of hydrohalite in frozen NaCl solutions by X-ray absorption spectroscopy


en: urbanization switzerland land-use modeling landscape-services statistical-model land-use-change

en: Data for Assessing potential landscape service trade-offs driven by urbanization in Switzerland

en:
The provided folder structure follows the set up of the scripts.
Data is only provided when it has been fully produced by us, otherwise dummies are given.
For data that will be produced by one of the scripts, no dummies are given.
The folders may contain sub-folders, which would be filled if the scripts were all run

 00_Analysis_Data
Data produced throughout the process of the calibration and projection that is later needed for analysis

 01_Raw_Data
Primary data from external sources that were used to compile the rasters and numbers used during the calibration and projection of landscape transitions as well as service calculation

 02_Prepared_Data
Data that will be prepared for further use by scripts are put in here
Additionally it contains the parameter sheets for the transition predictors that we compiled

 03_Model_Data
 Model outputs
Folder were the projection models statistics will be saved
 Models
Folder into which the actual transition models will be saved
 Residuals
Folder into which data on models residuals will be saved for future use

 04_Projected_Data
The output of the projections will be saved into this folder

 05_Service_Data
Any service calculations will be saved in here

 06_Results_Analysis
Any Analysis plots and tables as well as intermediate data will be saved here in the respective sub-folder

en: Scripts for Assessing potential landscape service trade-offs driven by urbanization in Switzerland

en:
Scripts ordered by their main purpose within sequential folders in this ReadMe marked with

Within the folders the capital letters refer to the parallel or sequential use of the scripts; scripts here in ReadMe are marked with

 00_Function Scripts
 Functions.r
Script containing several functions for analyzing statistical models: cohen.kappa, meva.table, D2, max-Tss, plot-TSS

 Service_functions.R
Script containing functions for calculating all services and returning sums (and median for bd and rec) as well as rasters

 01_Data Preparation
 A_AS_raster_preparation.R
Preparation of the land use rasters for the years 1985, 1997, and 2009 based on the original Arealstatistik (AS/ASCH); Assigning our categories and assining new values to all "construction sites" for calibrating the models
For model projection starting point (2009) and for service calculations (all ASCH years) the "other" categories were assigned to the land use categories based on dominant neighbor, so that there were no empty cells
Creation of water & mountainous area raster that stayed stable throughout the projections
Creation of MFH raster (2009), to have as base for all new projection years, as this category never becames any other

 B_Factors_preparation.R
Preparation of the factor rasters that were used to calculate service values per cell (see Table 2): LWfactor, Transportation, Vol, Rivers, Lakes, slope, TWW, Hazard_risk

 B_Predictors_preparation.R
Preparation of the raster of the predictors used for calibrating (ASCH years) and projecting (stable) the transition models that do not depend on the land use categories: Temperature (ASCH years), solar radiation (stable), Public transportation access (stable), Elevation (stable), Workers density (ASCH years), Population change (ASCH transitions)

 B_Regions_preparation.R
Preparation of bioregions as raster to be used to mask asxxxx later to calculated services and land use area just for one region

 C_Pred_proj.R
Projecting the dynamic predictors into the future for projecting the land use transitions with the model: Linear extrapolations for workers densities and population; more detailed extrapolation of temperature (based on data for all stations that matched between the temp table we used for the calibration period and the projections from CS2M, and then making a spatial interpolation for all years based on the interpolation of temperature for all years per station)

 02_Model calibration
 A_Bioreg_pred_corr.R
Calculating correlation between the predictors for each bioregion and land use category, Output are those with too high correlation so that one predictor can be removed from transitions were both predictors occur

 A_trans_numbers_bioreg.R
Calculate transition numbers for each bioregion to decide which transitions to primarily consider

 B_Bioreg_ext_eval.R
Calculating TSS based on the external evaluation for the bioregion transitions that happened often enough to try to model them (calibration data: 85-97, evaluation data: 97-09)

 B_Bioreg_int_eval.R
Calculating TSS based on the internal evaluation for the bioregion transitions that happened often enough to try to model them (calibration data: 70%, evaluation data: 30%)

 C_Bioreg_TSS_analysis.R
Extracting TSS numbers from internal and external evaluation, calculating mean and median for each transition for each region for each evaluation method and saving as csv per region to be able to select only the transitions with high enough model performance

 D_Bioreg_Models_for_prediction.R
Calibration of the models that were then used to project land use transitions into the future; calibrated separately for each bioregion

 E_Residuals_trans_models.R
Calculation of Moran's I statistics to analyze the residuals of the land use transition models

 03_Projections
 A_Bioreg_Projections_run.R
Master script for: Extrapolaitng into the future with the regional models and then assembling for each time step to calculate new AS predictors; Several Outputs: txt file to test for NAs in Predictors, txt file with random seed number for sampling, txt file with times for calculation steps (check progress of script while running), csv sheet with projected transition frequencies, tif-rasters for every region and time step, pdf with raster plot for every region and time step

 04_Service calculation
 A_Bioreg_as_services.R
Calculate services per bioregion for years 1985, 1997, 2009

 A_Service_calculation_bioregions.R
Extracting service numbers(sums) per bioregions for all projection runs and storing them in a list of matrixes to be available for further plotting; can be run for swiss wide projections or regional ones

 05_Analysis
 A_Bioreg_ASCH_development.R
Extracting the occurrence of each category per region for all runs (first part), then for ASCH years, and then plotting them nicely; only works for regional projections

 A_Bioreg_Predictors_importanceD2.R
Calculating D2 for each transition and each considered predictor from bivariate models to get value of importance

 B_Bioreg_Pred_imp_analysis.R
Using calculated D2 to sort Predictors according to their importance for each transition

 A_Bioreg_Spatial_results_analysis.R
Plotting randomly sampled runs for ASCH and services to analyses their spatial development

 A_Bioreg_Transition_numbers_check_v1.R
Analyzing the frequency for all transitions for all regions and producing the point plots for each transition separately

 A_Bioreg_Transition_numbers_check_v2.R
Analyzing percentage and numbers of all transitions but plotting them per region and originating ASCH category

 B_Transition_Analysis.R
Further Analysis of transition numbers

 A_Service_analysis.R
Plotting of the service development for bioregions in several ways for analyzing

en:
Data and scripts of publication:

Madleina Gerecke, Oskar Hagen, Janine Bolliger, Anna M. Hersperger, Felix Kienast, Bronwyn Price, Loïc Pellissier (2019) Assessing potential landscape service trade-offs driven by urbanization in Switzerland. Palgrave communications.

Contains land use projections for Switzerland and scripts and data for these projections as well as the calculation of landscape services.

Data Folder:
Contains sub-folder with the data necessary for this study (provided were no copyright issues, otherwise placeholders with descriptions), and folders where produced data may be stored

Scripts Folder:
Contains scripts organized into subfolders depending on their purpose

Note: Some abbreviations within the scripts and data are derived from German words and not English.

land-use

display_name: land-use

land-use-change

display_name: land-use-change

landscape-services

display_name: landscape-services

modeling

display_name: modeling

statistical-model

display_name: statistical-model

switzerland

display_name: switzerland

urbanization

display_name: urbanization

educ

en: Education, culture and sport

envidat

en: EnviDat: the Environmental Data Portal

en: EnviDat is the WSL data portal providing unified and managed access to environmental monitoring and research data. The portal has the capability to host and publish data sets. Through the sharing of unique and high-value data sets worldwide, EnviDat fosters research and contributes to advancing environmental science.

en: Land use projections and services for Switzerland


maintainer: Anita Risch

en: ecosystem-coupling ecosystem-multifunctionality grassland swiss-national-park ecosystem-functions vertebrates subalpine grazing exclusion invertebrates

en: Datafile

en:
ID: Plot ID composited of Fence & Treatment number (90 plots total)

Grassland: Name of the six subalpine grasslands where the exclosure setups were erected. In large grasslands (GRIM, MING, STAB), we established four hierarchical exclosure setups (two in short- and two in tall-grass vegetation), whereas in the small grasslands (BOTS, DADA, MARG) only one setup was erected per vegetation type. BOTS = Val dal Botsch, GRIM = Grimmels, MING = Val Mingér, STAB = Alp Stabelchod, DADA = Stabelchod da daint, MARG = Margunet

Fence: Number of exclosure setup (1 - 18). Note that uneven numbers (1,3,5,7,9,11,13,15,17) represent setups established on short-grass vegetation, even numbers (2,4,6,8,10,12,14,16,18) setups established in tall-grass vegetation
Vegetation	Vegetation type, 1 = short-grass vegetation, 2 = tall-grass vegetation; (9 replicate fences on each vegetation type)
Treatment	Progressive animal exclusion, numbers indicate which animals had access to the plot: 1 = Large/medium/small mammals, and invertebrates have access ("L/M/S/I"), 2 = Medium/small mammals, and invertebrates have access (“M/S/I”), 3 = Small mammals and invertebrates have access (“S/I”), 4 = Invertebrates have access (“I”), 5 = No animals have access (“None”)

Bulkden:	Fine fraction soil bulk density (g cm-3)

Soil.moist	: Soil moisture (%)

Soil.org.C: Rhizosphere soil C content (%)

pH: Soil pH

Nema.PC1: First principal component (PC1) of principal component analysis (PCA) of the soil nematode community composition

Nema.PC2: Second principal component (PC2) of principal component analysis (PCA) of the soil nematode community composition

Micro.PC1: 	First principal component (PC1) of principal component analysis (PCA) of the soil microbial community composition

Micro.PC2: Second principal component (PC2) of principal component analysis (PCA) of the soil microbial community composition

Abv.Inv.PC1: First principal component (PC1) of principal component analysis (PCA) of the aboveground-dwelling invertebrate community composition

Abv.Inv.PC2: Second principal component (PC2) of principal component analysis (PCA) of the aboveground-dwelling invertebrate community composition

Arthr.PC1: First principal component (PC1) of principal component analysis (PCA) of the soil arthropod community composition

Arthr.PC2: Second principal component (PC2) of principal component analysis (PCA) of the soil arthropod community composition

Plant.PC1: First principal component (PC1) of principal component analysis (PCA) of the plant community composition

Plant.PC2: Second principal component (PC2) of principal component analysis (PCA) of the plant community composition

Root.biomass: Root biomass (g m-2)

Micro.biomass	: Microbial biomass carbon (mg kg-1)

Plant.richness: Plant species richness

Plant.N: Plant tissue nitrogen content (%)

Soil.N.min: Soil net nitrogen mineralisation (kg ha-1 season)

Soil.resp: Soil carbon dioxide emissions (soil respiration; g m-2 h-1)

en:
This dataset contains all data on which the following publication below is based.

Paper Citation:
 > Risch AC, Ochoa-Hueso R, van der Putten WH, Bump JK, Busse MD, Frey B, Gwiazdowicz DJ, Page-Dumroese DS, Vandegehuchte ML, Zimmermann S, Schütz M. Size-dependent loss of aboveground animals differentially affects grassland ecosystem coupling and functions. 2018. Nature Communications 9: 3684. [doi: 10.1038/s41467-018-06105-4](https://doi.org/10.1038/s41467-018-06105-4).

Please cite this paper together with the citation for the datafile.

#Methods
Study sites
The experimental exclosure setups were installed within the SNP (IUCN category Ia preserve; Dudley 2008), in south-eastern Switzerland. The park covers 172 km2 of forests and subalpine and alpine grasslands along with scattered rock outcrops and scree slopes. The entire area has been protected from human impact (no hunting, fishing, camping or off-trail hiking) since 1914. Large, fairly homogenous patches of short- and tall-grass vegetation, which originate from different historical management and grazing regimes, cover the park’s subalpine grasslands entirely. Short-grass vegetation developed in areas where cattle used to rest (nutrient input) prior to the park’s foundation (14th century to 1914) (Schütz and others 2003, 2006) and is dominated by lawn grass species such as Festuca rubra L., Briza media L. and Agrostis capillaris L. (Schütz and others 2003, 2006). Today, this vegetation type is intensively grazed by diverse vertebrate and invertebrate communities that inhabit the park and consume up to 60% of the available biomass (Risch and others 2013). Tall-grass vegetation developed where cattle formerly grazed, but did not rest, and is dominated by rather nutrient-poor tussocks of Carex sempervirens Vill. and Nardus stricta L. (Schütz and others 2003, 2006). This vegetation type receives considerably less grazing, with only roughly 20% of the biomass consumed (Risch and others 2013). Consequently, the two vegetation types together represent a long-term trajectory of changes in grazing regimes. Underlying bedrock of all grasslands is dolomite, which renders these grasslands rather poor in nutrients regardless of former and current land-use regimes.
Experimental design
To progressively exclude aboveground vertebrate and invertebrate animals, we established 18 size-selective exclosure setups (nine in short-grass, nine in tall-grass vegetation) distributed over six subalpine grasslands across the SNP (Risch and others 2013, 2015). Elevation differences of exclosure locations did not exceed 350 m (between 1975 and 2300 m a.s.l.). The exclosures were established immediately after snowmelt in spring 2009 and were left in place for five consecutive growing seasons (until end of 2013). They were, however, temporarily dismantled every fall (late October after first snowfall) to protect them from avalanches. They were re-established in the same location every spring immediately after snowmelt. Each size-selective exclosure setup consisted of five plots (2 x 3 m) that progressively excluded aboveground vertebrates and invertebrates from large to small. The plots are labelled according to the guilds that had access to them “L/M/S/I”, “M/S/I”, “S/I”, “I”, “None”; L = large mammals, M = medium mammals, S = small mammals, I = invertebrates, None = no animals had access. As we only had permission to have the experimental setup in place for five consecutive growing seasons, the experiment had to be completely dismantled in the late fall of 2013 and all material removed from the SNP.
Our exclosure design was aimed at excluding mammalian herbivores, but naturally also excluded the few medium and small mammalian predators, as well as the entire aboveground invertebrate food web. A total of 26 large to small mammal species can be found in the SNP, but large apex predators are missing (wolf, bear, lynx) . Reptiles, amphibians and birds are scarce to absent in the subalpine grasslands under study. Only two reptile species occur in the park and they are confined to rocky areas that warm up enough for them to survive. One frog species spawns in an isolated pond far from our grasslands. Only three bird species occasionally feed on the subalpine grasslands. Using game cameras (Moultrie 6MP Game Spy I-60 Infrared Digital Game Camera, Moultrie Feeders, Alabaster, AL, USA), we did observe that the medium- and small-sized mammals (marmot/hares and mice) were not afraid to enter the fences and feed on their designated plots. We never spotted reptiles, amphibians or birds on camera. We distinguished between 59 higher aboveground-dwelling invertebrate taxa that our size-selective exclosures excluded (see also methods for aboveground-dwelling invertebrates below).
The “L/M/S/I” plot (not fenced) was located at least 5 m from the 2.1 m tall and 7 x 9 m large main electrical fence that enclosed the other four plots. The bottom wire of this fence was mounted at 0.5 m height and was not electrified to enable safe access for medium and small mammals, while fencing out the large ones. Within each main fence, we randomly established four 2 x 3 m plots separated by 1-m wide walkways from one another and from the main fence line: 1) the “M/S/I” plots were unfenced, allowing access to all but the large mammals; 2) the “S/I” plots (10 x 10 cm electrical mesh fence) excluded all medium-sized mammals. Note that the bottom 10 cm of this fence remained non-electrified to enable safe access for small mammals; 3) the “I” plots (2 x 2 cm metal mesh fence) excluded all mammals. We double-folded the mesh at the bottom 50 cm to reduce the mesh size to smaller than 1 x 1 cm openings; and 4) the “None” plots were surrounded by a 1 m tall mosquito net (1.5 x 2 mm) to exclude all animals. The top of the plot was covered with a mosquito-meshed wooden frame mounted to the corner posts (roof). We treated these plots a few times with biocompatible insecticide (Clean kill original, Eco Belle GmbH, Waldshut-Tiengen, Germany) to remove insects that might have entered during data collection or that hatched from the soil, but amounts were negligible and did not impact soil moisture conditions within these plots.
To assess whether the design of the “None” exclosure (mesh and roof) affected the response variables within the plots and, therefore, influenced the results, we established an additional six “micro-climate control” exclosures (one in each of the six grasslands) (Risch and others 2013, 2015). These exclosures were built as the “None” exclosures but were open at the bottom (20 cm) of the 3-m side of the fence facing away from the prevailing wind direction to allow invertebrates to enter. A 20-cm high and 3-m long strip of metal mesh was used to block access to small mammals. Thus, this construction allowed a comparable micro-climate to the “None” plots, but also a comparable feeding pressure by invertebrates to the “I” plots. We compared various properties within these exclosures against one another to assess if our construction altered the conditions in the “None” plots. We showed that differences in plant (e.g., vegetation height, aboveground biomass) and soil properties (e.g., soil temperature, moisture) found between the “I” and the “None” treatments were not due to the construction of the “None” exclosure, but a function of animal exclusions, although the amount of UV light reaching the plant canopy was significantly reduced (Risch and others 2013).
Aboveground invertebrate sampling
Aboveground invertebrates were sampled with two different methods to capture both ground- and plant-dwelling organisms: 1) we randomly placed two pitfall traps (67 mm in diameter, covered with a roof) filled with 20% propylene glycol in one 1 x 1 m subplot of the 2 x 3 m treatment plots in spring 2013 (May) and emptied them every two weeks until late September 2013 (Vandegehuchte and others 2017b, 2018). A pitfall trap consisted of a plastic cylinder (13 cm depth, 6.75 cm diameter). Within each cylinder we placed a 100 ml plastic vial with outer diameter 6.70 cm and on top of the cylinder we placed a plastic funnel to guide the invertebrates into the vials. Each trap was cover with a cone-shaped and transparent plastic roof to protect the trap from rain (Vandegehuchte and others 2017b, 2018). Note that in the “None” plots only one trap was placed as control to check for effectiveness of the exclosure. 2) We vacuumed all invertebrates from a 60 x 60 cm area on another 1 x 1 m subplot with a suction sampler (Vortis, Burkhard manufacturing CO, Ltd., Rickmansworth, Hertfordshire, UK) every month from June to September 2013 (Vandegehuchte and others 2017b, 2018). For this purpose, we quickly placed a square plastic frame (60 x 60 x 40 cm) with a closable mosquito mesh sleeve attached to the top edge into the plot from the outside. The suction sample was then inserted into through the sleeve and operated for 45 s to collect the invertebrates (Vandegehuchte and others 2017b, 2018).
We sorted the ≈100 000 individuals collected with both methods by hand and identified each individual morphologically to the lowest taxonomic level feasible (59 taxa, including orders, suborders, subfamilies, families; phylum for Mollusca). These taxa belonged to the following feeding types: 19 herbivores, 16 detritivores, 9 predators, 8 mixed feeders, 5 omnivores and 2 non-classified feeders (or not feeding as adults) (Vandegehuchte and others 2017b). We summed the numbers from the two pitfall traps and the suction sampling over the course of the 2013 season to represent the aboveground invertebrate abundance and community composition of a plot. Note: we did not specifically attempt to catch flying invertebrates with e.g., sticky traps, thus a few flying insects may have been missed with our vacuum sampling approach.
Sampling of plant properties
The vascular plant species composition was assessed at peak biomass every summer (July) by estimating the frequency of occurrence of each species with the pin count method in each plot (Frank and McNaughton 1990). A total of 172 taxa occurred within our 90 plots and we calculated plant species richness for each plot separately. We used the 2013 data in this study. Plant quality was assessed every year in July and September; here we use plant quality at the end of the experiment (September 2013). Two 10 x 100 cm wide strips of vegetation per plot were clipped, combined, dried at 65°C, and ground (Pulverisette 16, Fritsch, Idar-Oberstein, Germany) to pass through a 0.5 mm sieve. Twenty randomly selected samples across all treatments were analysed for N (Leco TruSpec Analyser, Leco, St. Joseph, Michigan, USA) (Vandegehuchte and others 2015). Nitrogen concentrations of the other samples were then estimated from models established for the experiment and the entire SNP relating Fourier transform-near infrared reflectance (FT-NIR) spectra to the measured values of N using a multi-purpose FT-NIR spectrometer (Bruker Optics, Fällanden, Switzerland) (Vandegehuchte and others 2015). Root biomass was sampled every fall by collecting five 2.2 cm diameter x 10 cm deep soil samples (Giddings Machine Company, Windsor, CO, USA) per plot (450 samples year-1). The samples were dried at 30 °C and roots were sorted from the sample by hand. We sorted each sample for 1 h which allowed to retrieve over 90% of all roots present in the samples (Risch and others 2013). The roots were then dried at 65 °C for 48 and weighed to the nearest mg. We averaged the values per plot and used the 2013 data only in this study.
Sampling of edaphic communities
In 2009, 2010, and 2011 we collected three composited soil samples (5 cm diameter x 10 cm depth; AMS Samplers, American Falls, ID, USA) and assessed bacterial community structure using T-RFLP profiling (Liu and others 1997; Blackwood and others 2003; Hodel and others 2014). We detected a total of 89 operational taxonomic units (OTUs). These values are in accordance with other studies reporting OTU richness (Wirthner and others 2011; Zumsteg and others 2012; Meola and others 2014) using T-RFLP profiling, a method that detects the most abundant, and thus likely, the most relevant, taxa. We averaged the data over the three years of collections for our calculations. Microbial biomass carbon (MBC) was determined with the substrate-induced method (Anderson and Domsch 1978) every fall (September) between 2009 and 2013 by collecting three mineral soil samples (5 cm diameter × 10 cm mineral soil core, AMS Samplers, American Falls, ID, USA). The three samples were combined (90 samples for each sampling year), immediately put on ice, taken to the laboratory, passed through a 2-mm sieve and stored at 4°C. Again, we only used the 2013 data in this study.
Soil samples (5 cm diameter x 10 cm depth) to extract soil arthropods were collected in June, July, and August 2011 with a soil corer lined with a plastic sleeve to ensure an undisturbed sample (total of 270 samples). The plastic line core was immediately sealed on both ends using cling film and put into a cooler. All plots were sampled within three days and the extraction of arthropods started the evening of the sampling day using a high-gradient Tullgren funnel apparatus (Crossley and Blair 1991; Vandegehuchte and others 2015). Samples were kept in the extractor for four days and the soil arthropods were collected in 95% ethanol. All individuals were counted and each individual was identified morphologically to the lowest level feasible [76 taxa, including orders, suborders, subfamilies, families (Protura, Thysanoptera, Aphidina, Psylina, Coleoptera, Brachycera, Nematocera, Auchenorryncha, Heteroptera, Formicidae); sub-phylum for Myriapoda, for Acari and Collembola also including morpho-species). Note that we also included larval stages (nine of the 76 taxa) (Vandegehuchte and others 2015). All data were summed over the season. A detailed species list for mites and collembolans is published (Vandegehuchte and others 2017a) [https://doi.org/10.1371/journal.pone.0118679.s001]. Earthworms are rare in the SNP and therefore were not included. We collected eight random 2.2 cm diameter x 10 cm deep soil cores from each plot in September 2013 to determine the soil nematode community composition. The samples were mixed and the nematodes were extracted from 100 ml of fresh soil using Oostenbrink elutriators (Oostenbrink 1960). All nematodes in a 1 ml of the 10 ml extract were counted, a minimum of 150 individuals sample-1 were identified to genus or family level using (Bongers 1988), the numbers of all nematodes were extrapolated to the entire sample and expressed for a 100 g dry sample. In total we identified 63 genus or family levels (Vandegehuchte and others 2015). The list of all the nematodes found is published (Vandegehuchte and others 2015) [http://www.oikosjournal.org/appendix/oik-03341] or DOI: [doi: 10.1111/oik.03341].
We are aware that sampling soil microbes from 2009 to 2011 and soil arthropods in 2011 was not ideal, but we are positive that this does not bias the results. Most of the parameters measured in our experiment either already showed a treatment response after the first growing season (e.g., plant biomass) or did not respond over the entire time experiment (e.g., microbial biomass C). The microbial community composition (2009 – 2011) was highly influenced by inter-annual differences in temperature and precipitation, but did not differ between treatments or vegetation types (Hodel and others 2014). We therefore felt comfortable using the 2009 through 2011 data for describing the soil microbial community in our experimental treatments. Similarly, we are positive that our soil arthropod data are representative. We did assess soil arthropods in August 2012 and found no differences to the August 2011 data. However, we did not feel comfortable combining the 2011 June, July, August data with only August data for 2012 for our analyses.
Sampling of soil properties
We collected three soil samples (5 cm diameter x 10 cm depth) in each plot in September 2013 after removing the vegetation. First, we collected the top layer of mineral soil rich in organic matter, the surface organic layer or rhizosphere, typically 1 to 3 cm in depth with a soil corer (AMS Samples, American Falls, Idaho, USA). Second, we collected a 10 cm mineral soil core beneath this surface layer. The cores for each layer were composited, dried at 65 °C for 48 h and fine-ground to pass a 0.5 mm screen. We then analysed all samples for total C using a Leco TruSpec Analyser (Leco, St. Joseph, Michigan, USA). Mineral soil pH was measured potentiometrically in 1:2 soil:CaCl2 solution with an equilibration time of 30 min.
Soil net N mineralisation was assessed during the 2013 growing season (Risch and others 2015). For this purpose, we randomly collected a 5 cm diameter x10 cm deep soil sample with a soil corer (AMS Samples, American Falls, Idaho, USA) after clipping the vegetation in June 2013. After weighing and sieving (4 mm mesh) the soil, we extracted a 20 g subsample in 1 mol l-1 KCl for 1.5 h on an end-over-end shaker and thereafter filtered it through ashless folded filter paper (DF 5895 150, ALBET LabScience, Hahnenmühle FineArt GmbH, Dassel, Germany). From these filtrates NO3- concentrations were measured colorimetrically (Norman and Stucki 1981) and NH4+with flow injection analysis (FIAS 300, Perkin Elmer, Waltham Massachusetts, USA) (Risch and others 2015). We dried the rest of the sample 105 °C to constant mass to determine fine,fraction bulk density. A second soil sample was collected within each plot in June 2013 with a corer lined with a 5 x 13 cm aluminium cylinder. The corer was driven 11.5 cm deep into the soil so that the top 1.5 cm of the cylinder remained empty. Into this space we placed a polyester bag (250 µm) filled an ion-exchanger resin to capture the incoming N. The bag was filled with a 1:1 mixture of acidic and alkaline exchanger resin (ion-exchanger I KA/ion exchanger IIIAA, Merck AG, Darmstadt, Germany). We then removed 1.5 cm soil at the bottom of the cylinder and placed a second resin exchanger bag into this space to capture the N leached from the soil column. To assure that the exchange resin was saturated with H+ and Cl- prior to filling the bags, the mixture was stirred with 1.2 ml l-1 HCl for 1 h and then rinsed with demineralized water until the electrical conductivity of the water reached 5 µm cm-1. The cylinder with the resin bags in place was reinserted into the soil with the top flush to the soil surface and incubated for three months. We recollected the cylinders in September 2013. Each resin bag and 20 g of sieved soil (4 mm mesh) from each cylinder were then separately extracted with KCl and NO3- and NH4+ concentrations were measured. Nitrate and NH4+ concentrations of all samples were then converted to a content basis by multiplying their values with fine fraction bulk density. Net N mineralisation was thereafter calculated as the difference between the N content of the samples collected at the end of the three-month incubation (including the N extracted from the bottom resin bag) and the N content at the beginning of the incubation (Risch and others 2015).
Soil CO2 emissions were measured every two weeks between 0900 and 1700 hrs from early May through late September 2013 with a PP-Systems SRC-1 soil respiration chamber (15 cm high, 10 cm diameter; closed circuit) attached to a PP-Systems EGM-4 infrared gas analyser (PP-Systems, Amesbury, MA, USA) on two locations per plot (Risch and others 2013). The chamber was placed on randomly placed, permanently installed PVC collars (10 cm diameter) driven 5 cm into the soil at the beginning of the study (Risch and others 2013). Freshly germinated plants growing within the collars were removed prior to each measurement to avoid measuring plant respiration or photosynthesis. The two measurements collected per plot and sampling date were averaged.
Soil moisture (with time domain reflectometry; Field-Scout TDR-100, Spectrum Technologies, Plainfield, Illionois, USA) and temperature (with a waterproof digital pocket thermometer; Barnstead International, Dubuque, Iowa, USA) were measured at five random locations per plot every two weeks during the growing seasons during the experiment for the 0 to 10 cm depth (Risch and others 2013, 2015). As soil moisture and soil temperature were highly negatively correlated (Risch and others 2013), we only used soil moisture for this study. We used plot-level averages of all values available to capture soil moisture variability during the five years of the experiment. The results remained unchanged when we only used soil moisture from the 2013 growing season.
Numeral calculations and statistical analyses
Ecosystem coupling. We conducted principal component analyses (PCAs; unscaled) at the complete dataset level using the abundances of each taxonomical entity to describe each of the five different communities used in this study: aboveground-dwelling invertebrates, vascular plants, soil microorganisms, soil arthropods and soil nematodes. We retained the first two components (PCA axis 1 and PCA axis 2) of each analysis as we found them to adequately represent the temporal and spatial variability of our 90 treatment plots in previous studies55,67. Together they explained a total of 71.70% of the variation for aboveground invertebrates, 44.36% for plants, 44.85% for soil microorganisms, 61.85% for soil arthropods and 77.19% for soil nematodes. In addition, we used soil pH and soil organic C content as a proxy for soil chemical properties, soil bulk density as a proxy for soil physical properties and soil moisture (negatively correlated with soil temperature) as a proxy for soil micro-climatic conditions for an overall total of fourteen constituents.
We calculated ecosystem coupling9 for each exclosure treatment within each vegetation type (i.e., 2  5 treatment combinations in total) as an integrated measure of pairwise ecological interactions between ecosystem constituents representing ecological communities and the soil abiotic environment. These ecological interactions are defined by non-parametric Spearman rank correlation analyses between two constituents, excluding interactions involving two abiotic constituents (e.g., soil pH vs. soil moisture) and interactions between the first (PC1) and second (PC2) component of each community type, as these are orthogonal by definition. Interactions between abiotic constituents were excluded from the analyses because the focus of our study was on communities and how they interact with one another and their surrounding environment; therefore, including abiotic-abiotic interactions was not of interest here. Given that the effectiveness of our experimental design resulted in that no community composition data of aboveground-dwelling invertebrates was available for the “None” plots (all animals excluded), only thirteen instead of fourteen constituents were included in the ecosystem coupling calculations for this treatment. The complete absence of aboveground invertebrates represents the most extreme case of disturbance between aboveground animal communities and the rest of the ecosystem constituents. This may have resulted in a slight overestimation of ecosystem coupling for these plots.
	Average ecosystem coupling was calculated as follows:
Ecosystem coupling=
where Xi is the absolute Coupling was calculated value of the Spearman’s rho coefficient of the ith correlation for each treatment within each vegetation type (i.e., based on nine replicates each), considering and n is the number of pairwise comparisons considered (n = a total of 80; interactions (56 in the case of the “None” treatment). We considered a total of 40 biotic-biotic interactions (i.e., concerning two community-level principal components such as plants and microbes; 24 in the case of the “None” treatment) and 40 abiotic-biotic (i.e., concerning one community-level principal component and one abiotic factor, e.g., plant community and soil properties; 32 in the case of the “None” treatment).	Coupling was calculated for each treatment within each vegetation type (i.e., based on nine replicates each), considering a total of 80 interactions (56 in the case of the “None” treatment). We considered a total of 40 biotic-biotic interactions (i.e., concerning two community-level principal components such as plants and microbes; 24 in the case of the “None” treatment) and 40 abiotic-biotic (i.e., concerning one community-level principal component and one abiotic factor, e.g., plant community and soil properties; 32 in the case of the “None” treatment).
To establish whether constituents were significantly and positively coupled within treatments (i.e., the average of their correlation coefficients were greater than in a null model where correlation only happens by chance), we calculated one-tailed p-values based on permutation tests with 999 permutations.

We considered six ecosystem functions and process rates commonly used to assess ecosystem functioning (Meyer and others 2015; Manning and others 2018). Plant N content represents a measure of forage quality, while plant richness has been shown to stabilise biomass production, thus allowing the system to respond to changes in herbivory. Soil net N mineralisation, soil respiration, root biomass, and microbial biomass represent fluxes or stocks of energy. For all functions and processes higher values represent higher functioning (Manning and others 2018). All these variables were measured in the last year of the experiment (2013). We then quantified ecosystem multifunctionality using the multiple threshold approach (Byrnes and others 2014; Manning and others 2018), which considers the number of functions that are above a certain threshold, over a series of threshold values (typically 10-99%) that are defined based on the maximum value of each function. We weighted all our functions equally for these calculations (Manning and others 2018). The number of functions in a plot with values higher than a given threshold value for the respective function is summed up. The sum represents ecosystem multifunctionality for that plot. Given that choosing any particular threshold as a measure of ecosystem multifunctionality is arbitrary, we calculated the average of thresholds from 10-90% (in 10% intervals) as a more integrated representation of ecosystem multifunctionality.
We used Pearson correlations to explore the relationships between ecosystem coupling (all interactions, biotic-biotic interactions, abiotic-biotic interactions involving above- and belowground constituents, and all interactions, biotic-biotic interactions, abiotic-biotic interactions involving belowground constituents only) and ecosystem multifunctionality by calculating the slopes of all relationships between ecosystem coupling and multifunctionality for all thresholds between 10 and 99%. We also related ecosystem coupling with the average of multifunctionality at thresholds between 30-80% as explained before and considered this correlation as a robust indication of the type of association between these two variables. In addition, we explored the relationships between ecosystem coupling (all interactions, biotic-biotic interactions, abiotic-biotic interactions involving above- and belowground constituents, and all interactions, biotic-biotic interactions, abiotic-biotic interactions involving belowground constituents only) and individual ecosystem functions. The effects of exclosures and vegetation type on individual functions and multifunctionality were evaluated using linear mixed effects models ('lme' function of the nlme package), with exclosure and vegetation type as fixed effects and fence as a random factor. All statistical analyses and numerical calculations were done in R version 3.4.0 (R Core Team 2016).

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ecosystem-coupling

display_name: ecosystem-coupling

ecosystem-functions

display_name: ecosystem-functions

ecosystem-multifunctionality

display_name: ecosystem-multifunctionality

exclusion

display_name: exclusion

grassland

display_name: grassland

grazing

display_name: grazing

invertebrates

display_name: invertebrates

subalpine

display_name: subalpine

swiss-national-park

display_name: swiss-national-park

vertebrates

display_name: vertebrates

en: Ecosystem coupling and multifunctionality - exclosure experiment