maintainer: Anita Risch

en: invertebrates vertebrates grazing ecosystem-coupling exclusion swiss-national-park subalpine ecosystem-functions grassland ecosystem-multifunctionality

en: Datafile

en:
ID: Plot ID composited of Fence & Treatment number (90 plots total)

Grassland: Name of the six subalpine grasslands where the exclosure setups were erected. In large grasslands (GRIM, MING, STAB), we established four hierarchical exclosure setups (two in short- and two in tall-grass vegetation), whereas in the small grasslands (BOTS, DADA, MARG) only one setup was erected per vegetation type. BOTS = Val dal Botsch, GRIM = Grimmels, MING = Val Mingér, STAB = Alp Stabelchod, DADA = Stabelchod da daint, MARG = Margunet

Fence: Number of exclosure setup (1 - 18). Note that uneven numbers (1,3,5,7,9,11,13,15,17) represent setups established on short-grass vegetation, even numbers (2,4,6,8,10,12,14,16,18) setups established in tall-grass vegetation
Vegetation	Vegetation type, 1 = short-grass vegetation, 2 = tall-grass vegetation; (9 replicate fences on each vegetation type)
Treatment	Progressive animal exclusion, numbers indicate which animals had access to the plot: 1 = Large/medium/small mammals, and invertebrates have access ("L/M/S/I"), 2 = Medium/small mammals, and invertebrates have access (“M/S/I”), 3 = Small mammals and invertebrates have access (“S/I”), 4 = Invertebrates have access (“I”), 5 = No animals have access (“None”)

Bulkden:	Fine fraction soil bulk density (g cm-3)

Soil.moist	: Soil moisture (%)

Soil.org.C: Rhizosphere soil C content (%)

pH: Soil pH

Nema.PC1: First principal component (PC1) of principal component analysis (PCA) of the soil nematode community composition

Nema.PC2: Second principal component (PC2) of principal component analysis (PCA) of the soil nematode community composition

Micro.PC1: 	First principal component (PC1) of principal component analysis (PCA) of the soil microbial community composition

Micro.PC2: Second principal component (PC2) of principal component analysis (PCA) of the soil microbial community composition

Abv.Inv.PC1: First principal component (PC1) of principal component analysis (PCA) of the aboveground-dwelling invertebrate community composition

Abv.Inv.PC2: Second principal component (PC2) of principal component analysis (PCA) of the aboveground-dwelling invertebrate community composition

Arthr.PC1: First principal component (PC1) of principal component analysis (PCA) of the soil arthropod community composition

Arthr.PC2: Second principal component (PC2) of principal component analysis (PCA) of the soil arthropod community composition

Plant.PC1: First principal component (PC1) of principal component analysis (PCA) of the plant community composition

Plant.PC2: Second principal component (PC2) of principal component analysis (PCA) of the plant community composition

Root.biomass: Root biomass (g m-2)

Micro.biomass	: Microbial biomass carbon (mg kg-1)

Plant.richness: Plant species richness

Plant.N: Plant tissue nitrogen content (%)

Soil.N.min: Soil net nitrogen mineralisation (kg ha-1 season)

Soil.resp: Soil carbon dioxide emissions (soil respiration; g m-2 h-1)

en:
This dataset contains all data on which the following publication below is based.

Paper Citation:
 > Risch AC, Ochoa-Hueso R, van der Putten WH, Bump JK, Busse MD, Frey B, Gwiazdowicz DJ, Page-Dumroese DS, Vandegehuchte ML, Zimmermann S, Schütz M. Size-dependent loss of aboveground animals differentially affects grassland ecosystem coupling and functions. 2018. Nature Communications 9: 3684. [doi: 10.1038/s41467-018-06105-4](https://doi.org/10.1038/s41467-018-06105-4).

Please cite this paper together with the citation for the datafile.

#Methods
Study sites
The experimental exclosure setups were installed within the SNP (IUCN category Ia preserve; Dudley 2008), in south-eastern Switzerland. The park covers 172 km2 of forests and subalpine and alpine grasslands along with scattered rock outcrops and scree slopes. The entire area has been protected from human impact (no hunting, fishing, camping or off-trail hiking) since 1914. Large, fairly homogenous patches of short- and tall-grass vegetation, which originate from different historical management and grazing regimes, cover the park’s subalpine grasslands entirely. Short-grass vegetation developed in areas where cattle used to rest (nutrient input) prior to the park’s foundation (14th century to 1914) (Schütz and others 2003, 2006) and is dominated by lawn grass species such as Festuca rubra L., Briza media L. and Agrostis capillaris L. (Schütz and others 2003, 2006). Today, this vegetation type is intensively grazed by diverse vertebrate and invertebrate communities that inhabit the park and consume up to 60% of the available biomass (Risch and others 2013). Tall-grass vegetation developed where cattle formerly grazed, but did not rest, and is dominated by rather nutrient-poor tussocks of Carex sempervirens Vill. and Nardus stricta L. (Schütz and others 2003, 2006). This vegetation type receives considerably less grazing, with only roughly 20% of the biomass consumed (Risch and others 2013). Consequently, the two vegetation types together represent a long-term trajectory of changes in grazing regimes. Underlying bedrock of all grasslands is dolomite, which renders these grasslands rather poor in nutrients regardless of former and current land-use regimes.
Experimental design
To progressively exclude aboveground vertebrate and invertebrate animals, we established 18 size-selective exclosure setups (nine in short-grass, nine in tall-grass vegetation) distributed over six subalpine grasslands across the SNP (Risch and others 2013, 2015). Elevation differences of exclosure locations did not exceed 350 m (between 1975 and 2300 m a.s.l.). The exclosures were established immediately after snowmelt in spring 2009 and were left in place for five consecutive growing seasons (until end of 2013). They were, however, temporarily dismantled every fall (late October after first snowfall) to protect them from avalanches. They were re-established in the same location every spring immediately after snowmelt. Each size-selective exclosure setup consisted of five plots (2 x 3 m) that progressively excluded aboveground vertebrates and invertebrates from large to small. The plots are labelled according to the guilds that had access to them “L/M/S/I”, “M/S/I”, “S/I”, “I”, “None”; L = large mammals, M = medium mammals, S = small mammals, I = invertebrates, None = no animals had access. As we only had permission to have the experimental setup in place for five consecutive growing seasons, the experiment had to be completely dismantled in the late fall of 2013 and all material removed from the SNP.
Our exclosure design was aimed at excluding mammalian herbivores, but naturally also excluded the few medium and small mammalian predators, as well as the entire aboveground invertebrate food web. A total of 26 large to small mammal species can be found in the SNP, but large apex predators are missing (wolf, bear, lynx) . Reptiles, amphibians and birds are scarce to absent in the subalpine grasslands under study. Only two reptile species occur in the park and they are confined to rocky areas that warm up enough for them to survive. One frog species spawns in an isolated pond far from our grasslands. Only three bird species occasionally feed on the subalpine grasslands. Using game cameras (Moultrie 6MP Game Spy I-60 Infrared Digital Game Camera, Moultrie Feeders, Alabaster, AL, USA), we did observe that the medium- and small-sized mammals (marmot/hares and mice) were not afraid to enter the fences and feed on their designated plots. We never spotted reptiles, amphibians or birds on camera. We distinguished between 59 higher aboveground-dwelling invertebrate taxa that our size-selective exclosures excluded (see also methods for aboveground-dwelling invertebrates below).
The “L/M/S/I” plot (not fenced) was located at least 5 m from the 2.1 m tall and 7 x 9 m large main electrical fence that enclosed the other four plots. The bottom wire of this fence was mounted at 0.5 m height and was not electrified to enable safe access for medium and small mammals, while fencing out the large ones. Within each main fence, we randomly established four 2 x 3 m plots separated by 1-m wide walkways from one another and from the main fence line: 1) the “M/S/I” plots were unfenced, allowing access to all but the large mammals; 2) the “S/I” plots (10 x 10 cm electrical mesh fence) excluded all medium-sized mammals. Note that the bottom 10 cm of this fence remained non-electrified to enable safe access for small mammals; 3) the “I” plots (2 x 2 cm metal mesh fence) excluded all mammals. We double-folded the mesh at the bottom 50 cm to reduce the mesh size to smaller than 1 x 1 cm openings; and 4) the “None” plots were surrounded by a 1 m tall mosquito net (1.5 x 2 mm) to exclude all animals. The top of the plot was covered with a mosquito-meshed wooden frame mounted to the corner posts (roof). We treated these plots a few times with biocompatible insecticide (Clean kill original, Eco Belle GmbH, Waldshut-Tiengen, Germany) to remove insects that might have entered during data collection or that hatched from the soil, but amounts were negligible and did not impact soil moisture conditions within these plots.
To assess whether the design of the “None” exclosure (mesh and roof) affected the response variables within the plots and, therefore, influenced the results, we established an additional six “micro-climate control” exclosures (one in each of the six grasslands) (Risch and others 2013, 2015). These exclosures were built as the “None” exclosures but were open at the bottom (20 cm) of the 3-m side of the fence facing away from the prevailing wind direction to allow invertebrates to enter. A 20-cm high and 3-m long strip of metal mesh was used to block access to small mammals. Thus, this construction allowed a comparable micro-climate to the “None” plots, but also a comparable feeding pressure by invertebrates to the “I” plots. We compared various properties within these exclosures against one another to assess if our construction altered the conditions in the “None” plots. We showed that differences in plant (e.g., vegetation height, aboveground biomass) and soil properties (e.g., soil temperature, moisture) found between the “I” and the “None” treatments were not due to the construction of the “None” exclosure, but a function of animal exclusions, although the amount of UV light reaching the plant canopy was significantly reduced (Risch and others 2013).
Aboveground invertebrate sampling
Aboveground invertebrates were sampled with two different methods to capture both ground- and plant-dwelling organisms: 1) we randomly placed two pitfall traps (67 mm in diameter, covered with a roof) filled with 20% propylene glycol in one 1 x 1 m subplot of the 2 x 3 m treatment plots in spring 2013 (May) and emptied them every two weeks until late September 2013 (Vandegehuchte and others 2017b, 2018). A pitfall trap consisted of a plastic cylinder (13 cm depth, 6.75 cm diameter). Within each cylinder we placed a 100 ml plastic vial with outer diameter 6.70 cm and on top of the cylinder we placed a plastic funnel to guide the invertebrates into the vials. Each trap was cover with a cone-shaped and transparent plastic roof to protect the trap from rain (Vandegehuchte and others 2017b, 2018). Note that in the “None” plots only one trap was placed as control to check for effectiveness of the exclosure. 2) We vacuumed all invertebrates from a 60 x 60 cm area on another 1 x 1 m subplot with a suction sampler (Vortis, Burkhard manufacturing CO, Ltd., Rickmansworth, Hertfordshire, UK) every month from June to September 2013 (Vandegehuchte and others 2017b, 2018). For this purpose, we quickly placed a square plastic frame (60 x 60 x 40 cm) with a closable mosquito mesh sleeve attached to the top edge into the plot from the outside. The suction sample was then inserted into through the sleeve and operated for 45 s to collect the invertebrates (Vandegehuchte and others 2017b, 2018).
We sorted the ≈100 000 individuals collected with both methods by hand and identified each individual morphologically to the lowest taxonomic level feasible (59 taxa, including orders, suborders, subfamilies, families; phylum for Mollusca). These taxa belonged to the following feeding types: 19 herbivores, 16 detritivores, 9 predators, 8 mixed feeders, 5 omnivores and 2 non-classified feeders (or not feeding as adults) (Vandegehuchte and others 2017b). We summed the numbers from the two pitfall traps and the suction sampling over the course of the 2013 season to represent the aboveground invertebrate abundance and community composition of a plot. Note: we did not specifically attempt to catch flying invertebrates with e.g., sticky traps, thus a few flying insects may have been missed with our vacuum sampling approach.
Sampling of plant properties
The vascular plant species composition was assessed at peak biomass every summer (July) by estimating the frequency of occurrence of each species with the pin count method in each plot (Frank and McNaughton 1990). A total of 172 taxa occurred within our 90 plots and we calculated plant species richness for each plot separately. We used the 2013 data in this study. Plant quality was assessed every year in July and September; here we use plant quality at the end of the experiment (September 2013). Two 10 x 100 cm wide strips of vegetation per plot were clipped, combined, dried at 65°C, and ground (Pulverisette 16, Fritsch, Idar-Oberstein, Germany) to pass through a 0.5 mm sieve. Twenty randomly selected samples across all treatments were analysed for N (Leco TruSpec Analyser, Leco, St. Joseph, Michigan, USA) (Vandegehuchte and others 2015). Nitrogen concentrations of the other samples were then estimated from models established for the experiment and the entire SNP relating Fourier transform-near infrared reflectance (FT-NIR) spectra to the measured values of N using a multi-purpose FT-NIR spectrometer (Bruker Optics, Fällanden, Switzerland) (Vandegehuchte and others 2015). Root biomass was sampled every fall by collecting five 2.2 cm diameter x 10 cm deep soil samples (Giddings Machine Company, Windsor, CO, USA) per plot (450 samples year-1). The samples were dried at 30 °C and roots were sorted from the sample by hand. We sorted each sample for 1 h which allowed to retrieve over 90% of all roots present in the samples (Risch and others 2013). The roots were then dried at 65 °C for 48 and weighed to the nearest mg. We averaged the values per plot and used the 2013 data only in this study.
Sampling of edaphic communities
In 2009, 2010, and 2011 we collected three composited soil samples (5 cm diameter x 10 cm depth; AMS Samplers, American Falls, ID, USA) and assessed bacterial community structure using T-RFLP profiling (Liu and others 1997; Blackwood and others 2003; Hodel and others 2014). We detected a total of 89 operational taxonomic units (OTUs). These values are in accordance with other studies reporting OTU richness (Wirthner and others 2011; Zumsteg and others 2012; Meola and others 2014) using T-RFLP profiling, a method that detects the most abundant, and thus likely, the most relevant, taxa. We averaged the data over the three years of collections for our calculations. Microbial biomass carbon (MBC) was determined with the substrate-induced method (Anderson and Domsch 1978) every fall (September) between 2009 and 2013 by collecting three mineral soil samples (5 cm diameter × 10 cm mineral soil core, AMS Samplers, American Falls, ID, USA). The three samples were combined (90 samples for each sampling year), immediately put on ice, taken to the laboratory, passed through a 2-mm sieve and stored at 4°C. Again, we only used the 2013 data in this study.
Soil samples (5 cm diameter x 10 cm depth) to extract soil arthropods were collected in June, July, and August 2011 with a soil corer lined with a plastic sleeve to ensure an undisturbed sample (total of 270 samples). The plastic line core was immediately sealed on both ends using cling film and put into a cooler. All plots were sampled within three days and the extraction of arthropods started the evening of the sampling day using a high-gradient Tullgren funnel apparatus (Crossley and Blair 1991; Vandegehuchte and others 2015). Samples were kept in the extractor for four days and the soil arthropods were collected in 95% ethanol. All individuals were counted and each individual was identified morphologically to the lowest level feasible [76 taxa, including orders, suborders, subfamilies, families (Protura, Thysanoptera, Aphidina, Psylina, Coleoptera, Brachycera, Nematocera, Auchenorryncha, Heteroptera, Formicidae); sub-phylum for Myriapoda, for Acari and Collembola also including morpho-species). Note that we also included larval stages (nine of the 76 taxa) (Vandegehuchte and others 2015). All data were summed over the season. A detailed species list for mites and collembolans is published (Vandegehuchte and others 2017a) [https://doi.org/10.1371/journal.pone.0118679.s001]. Earthworms are rare in the SNP and therefore were not included. We collected eight random 2.2 cm diameter x 10 cm deep soil cores from each plot in September 2013 to determine the soil nematode community composition. The samples were mixed and the nematodes were extracted from 100 ml of fresh soil using Oostenbrink elutriators (Oostenbrink 1960). All nematodes in a 1 ml of the 10 ml extract were counted, a minimum of 150 individuals sample-1 were identified to genus or family level using (Bongers 1988), the numbers of all nematodes were extrapolated to the entire sample and expressed for a 100 g dry sample. In total we identified 63 genus or family levels (Vandegehuchte and others 2015). The list of all the nematodes found is published (Vandegehuchte and others 2015) [http://www.oikosjournal.org/appendix/oik-03341] or DOI: [doi: 10.1111/oik.03341].
We are aware that sampling soil microbes from 2009 to 2011 and soil arthropods in 2011 was not ideal, but we are positive that this does not bias the results. Most of the parameters measured in our experiment either already showed a treatment response after the first growing season (e.g., plant biomass) or did not respond over the entire time experiment (e.g., microbial biomass C). The microbial community composition (2009 – 2011) was highly influenced by inter-annual differences in temperature and precipitation, but did not differ between treatments or vegetation types (Hodel and others 2014). We therefore felt comfortable using the 2009 through 2011 data for describing the soil microbial community in our experimental treatments. Similarly, we are positive that our soil arthropod data are representative. We did assess soil arthropods in August 2012 and found no differences to the August 2011 data. However, we did not feel comfortable combining the 2011 June, July, August data with only August data for 2012 for our analyses.
Sampling of soil properties
We collected three soil samples (5 cm diameter x 10 cm depth) in each plot in September 2013 after removing the vegetation. First, we collected the top layer of mineral soil rich in organic matter, the surface organic layer or rhizosphere, typically 1 to 3 cm in depth with a soil corer (AMS Samples, American Falls, Idaho, USA). Second, we collected a 10 cm mineral soil core beneath this surface layer. The cores for each layer were composited, dried at 65 °C for 48 h and fine-ground to pass a 0.5 mm screen. We then analysed all samples for total C using a Leco TruSpec Analyser (Leco, St. Joseph, Michigan, USA). Mineral soil pH was measured potentiometrically in 1:2 soil:CaCl2 solution with an equilibration time of 30 min.
Soil net N mineralisation was assessed during the 2013 growing season (Risch and others 2015). For this purpose, we randomly collected a 5 cm diameter x10 cm deep soil sample with a soil corer (AMS Samples, American Falls, Idaho, USA) after clipping the vegetation in June 2013. After weighing and sieving (4 mm mesh) the soil, we extracted a 20 g subsample in 1 mol l-1 KCl for 1.5 h on an end-over-end shaker and thereafter filtered it through ashless folded filter paper (DF 5895 150, ALBET LabScience, Hahnenmühle FineArt GmbH, Dassel, Germany). From these filtrates NO3- concentrations were measured colorimetrically (Norman and Stucki 1981) and NH4+with flow injection analysis (FIAS 300, Perkin Elmer, Waltham Massachusetts, USA) (Risch and others 2015). We dried the rest of the sample 105 °C to constant mass to determine fine,fraction bulk density. A second soil sample was collected within each plot in June 2013 with a corer lined with a 5 x 13 cm aluminium cylinder. The corer was driven 11.5 cm deep into the soil so that the top 1.5 cm of the cylinder remained empty. Into this space we placed a polyester bag (250 µm) filled an ion-exchanger resin to capture the incoming N. The bag was filled with a 1:1 mixture of acidic and alkaline exchanger resin (ion-exchanger I KA/ion exchanger IIIAA, Merck AG, Darmstadt, Germany). We then removed 1.5 cm soil at the bottom of the cylinder and placed a second resin exchanger bag into this space to capture the N leached from the soil column. To assure that the exchange resin was saturated with H+ and Cl- prior to filling the bags, the mixture was stirred with 1.2 ml l-1 HCl for 1 h and then rinsed with demineralized water until the electrical conductivity of the water reached 5 µm cm-1. The cylinder with the resin bags in place was reinserted into the soil with the top flush to the soil surface and incubated for three months. We recollected the cylinders in September 2013. Each resin bag and 20 g of sieved soil (4 mm mesh) from each cylinder were then separately extracted with KCl and NO3- and NH4+ concentrations were measured. Nitrate and NH4+ concentrations of all samples were then converted to a content basis by multiplying their values with fine fraction bulk density. Net N mineralisation was thereafter calculated as the difference between the N content of the samples collected at the end of the three-month incubation (including the N extracted from the bottom resin bag) and the N content at the beginning of the incubation (Risch and others 2015).
Soil CO2 emissions were measured every two weeks between 0900 and 1700 hrs from early May through late September 2013 with a PP-Systems SRC-1 soil respiration chamber (15 cm high, 10 cm diameter; closed circuit) attached to a PP-Systems EGM-4 infrared gas analyser (PP-Systems, Amesbury, MA, USA) on two locations per plot (Risch and others 2013). The chamber was placed on randomly placed, permanently installed PVC collars (10 cm diameter) driven 5 cm into the soil at the beginning of the study (Risch and others 2013). Freshly germinated plants growing within the collars were removed prior to each measurement to avoid measuring plant respiration or photosynthesis. The two measurements collected per plot and sampling date were averaged.
Soil moisture (with time domain reflectometry; Field-Scout TDR-100, Spectrum Technologies, Plainfield, Illionois, USA) and temperature (with a waterproof digital pocket thermometer; Barnstead International, Dubuque, Iowa, USA) were measured at five random locations per plot every two weeks during the growing seasons during the experiment for the 0 to 10 cm depth (Risch and others 2013, 2015). As soil moisture and soil temperature were highly negatively correlated (Risch and others 2013), we only used soil moisture for this study. We used plot-level averages of all values available to capture soil moisture variability during the five years of the experiment. The results remained unchanged when we only used soil moisture from the 2013 growing season.
Numeral calculations and statistical analyses
Ecosystem coupling. We conducted principal component analyses (PCAs; unscaled) at the complete dataset level using the abundances of each taxonomical entity to describe each of the five different communities used in this study: aboveground-dwelling invertebrates, vascular plants, soil microorganisms, soil arthropods and soil nematodes. We retained the first two components (PCA axis 1 and PCA axis 2) of each analysis as we found them to adequately represent the temporal and spatial variability of our 90 treatment plots in previous studies55,67. Together they explained a total of 71.70% of the variation for aboveground invertebrates, 44.36% for plants, 44.85% for soil microorganisms, 61.85% for soil arthropods and 77.19% for soil nematodes. In addition, we used soil pH and soil organic C content as a proxy for soil chemical properties, soil bulk density as a proxy for soil physical properties and soil moisture (negatively correlated with soil temperature) as a proxy for soil micro-climatic conditions for an overall total of fourteen constituents.
We calculated ecosystem coupling9 for each exclosure treatment within each vegetation type (i.e., 2  5 treatment combinations in total) as an integrated measure of pairwise ecological interactions between ecosystem constituents representing ecological communities and the soil abiotic environment. These ecological interactions are defined by non-parametric Spearman rank correlation analyses between two constituents, excluding interactions involving two abiotic constituents (e.g., soil pH vs. soil moisture) and interactions between the first (PC1) and second (PC2) component of each community type, as these are orthogonal by definition. Interactions between abiotic constituents were excluded from the analyses because the focus of our study was on communities and how they interact with one another and their surrounding environment; therefore, including abiotic-abiotic interactions was not of interest here. Given that the effectiveness of our experimental design resulted in that no community composition data of aboveground-dwelling invertebrates was available for the “None” plots (all animals excluded), only thirteen instead of fourteen constituents were included in the ecosystem coupling calculations for this treatment. The complete absence of aboveground invertebrates represents the most extreme case of disturbance between aboveground animal communities and the rest of the ecosystem constituents. This may have resulted in a slight overestimation of ecosystem coupling for these plots.
	Average ecosystem coupling was calculated as follows:
Ecosystem coupling=
where Xi is the absolute Coupling was calculated value of the Spearman’s rho coefficient of the ith correlation for each treatment within each vegetation type (i.e., based on nine replicates each), considering and n is the number of pairwise comparisons considered (n = a total of 80; interactions (56 in the case of the “None” treatment). We considered a total of 40 biotic-biotic interactions (i.e., concerning two community-level principal components such as plants and microbes; 24 in the case of the “None” treatment) and 40 abiotic-biotic (i.e., concerning one community-level principal component and one abiotic factor, e.g., plant community and soil properties; 32 in the case of the “None” treatment).	Coupling was calculated for each treatment within each vegetation type (i.e., based on nine replicates each), considering a total of 80 interactions (56 in the case of the “None” treatment). We considered a total of 40 biotic-biotic interactions (i.e., concerning two community-level principal components such as plants and microbes; 24 in the case of the “None” treatment) and 40 abiotic-biotic (i.e., concerning one community-level principal component and one abiotic factor, e.g., plant community and soil properties; 32 in the case of the “None” treatment).
To establish whether constituents were significantly and positively coupled within treatments (i.e., the average of their correlation coefficients were greater than in a null model where correlation only happens by chance), we calculated one-tailed p-values based on permutation tests with 999 permutations.

We considered six ecosystem functions and process rates commonly used to assess ecosystem functioning (Meyer and others 2015; Manning and others 2018). Plant N content represents a measure of forage quality, while plant richness has been shown to stabilise biomass production, thus allowing the system to respond to changes in herbivory. Soil net N mineralisation, soil respiration, root biomass, and microbial biomass represent fluxes or stocks of energy. For all functions and processes higher values represent higher functioning (Manning and others 2018). All these variables were measured in the last year of the experiment (2013). We then quantified ecosystem multifunctionality using the multiple threshold approach (Byrnes and others 2014; Manning and others 2018), which considers the number of functions that are above a certain threshold, over a series of threshold values (typically 10-99%) that are defined based on the maximum value of each function. We weighted all our functions equally for these calculations (Manning and others 2018). The number of functions in a plot with values higher than a given threshold value for the respective function is summed up. The sum represents ecosystem multifunctionality for that plot. Given that choosing any particular threshold as a measure of ecosystem multifunctionality is arbitrary, we calculated the average of thresholds from 10-90% (in 10% intervals) as a more integrated representation of ecosystem multifunctionality.
We used Pearson correlations to explore the relationships between ecosystem coupling (all interactions, biotic-biotic interactions, abiotic-biotic interactions involving above- and belowground constituents, and all interactions, biotic-biotic interactions, abiotic-biotic interactions involving belowground constituents only) and ecosystem multifunctionality by calculating the slopes of all relationships between ecosystem coupling and multifunctionality for all thresholds between 10 and 99%. We also related ecosystem coupling with the average of multifunctionality at thresholds between 30-80% as explained before and considered this correlation as a robust indication of the type of association between these two variables. In addition, we explored the relationships between ecosystem coupling (all interactions, biotic-biotic interactions, abiotic-biotic interactions involving above- and belowground constituents, and all interactions, biotic-biotic interactions, abiotic-biotic interactions involving belowground constituents only) and individual ecosystem functions. The effects of exclosures and vegetation type on individual functions and multifunctionality were evaluated using linear mixed effects models ('lme' function of the nlme package), with exclosure and vegetation type as fixed effects and fence as a random factor. All statistical analyses and numerical calculations were done in R version 3.4.0 (R Core Team 2016).

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- R Core Team. 2016. R: A language and environment for statistical computing. Vienna, Austria: R Foundation for Statistical Computing
- Risch AC, Haynes AG, Busse MD, Filli F, Schütz M. 2013. The response of soil CO2 fluxes to progressively excluding vertebrate and invertebrate herbivores depends on ecosystem type. Ecosystems 16:1192–202.
- Risch AC, Schütz M, Vandegehuchte ML, Van Der Putten WH, Duyts H, Raschein U, Gwiazdowicz DJ, Busse MD, Page-Dumroese DS, Zimmermann S. 2015. Aboveground vertebrate and invertebrate herbivore impact on net N mineralization in subalpine grasslands. Ecology 96:3312–22.
- Schütz M, Risch AC, Achermann G, Thiel-Egenter C, Page-Dumroese DS, Jurgensen MF, Edwards PJ. 2006. Phosphorus translocation by red deer on a subalpine grassland in the Central European Alps. Ecosystems 9:624–633.
- Schütz M, Risch AC, Leuzinger E, Krüsi BO, Achermann G. 2003. Impact of herbivory by red deer (Cervus elaphus L.) on patterns and processes in subalpine grasslands in the Swiss National Park. For Ecol Manage 181:177–88.
- Vandegehuchte ML, van der Putten WH, Duyts H, Schütz M, Risch AC. 2017a. Aboveground mammal and invertebrate exclusions cause consistent changes in soil food webs of two subalpine grassland types, but mechanisms are system-speciﬁc. Oikos 126:212–23.
- Vandegehuchte ML, Raschein U, Schütz M, Gwiazdowicz DJ, Risch AC. 2015. Indirect short- and long-term effects of aboveground invertebrate and vertebrate herbivores on soil microarthropod communities. PLoS One 10:e0118679.
- Vandegehuchte ML, Schütz M, de Schaetzen F, Risch AC. 2017b. Mammal-induced trophic cascades in invertebrate food webs are modulated by grazing intensity in subalpine grassland. J Anim Ecol 86:1434–46.
- Vandegehuchte ML, Trivellone V, Schütz M, Firn J, de Schaetzen F, Risch AC. 2018. Mammalian herbivores affect leafhoppers associated with specific plant functional types at different timescales. Funct Ecol 32:545–55.
- Wirthner S, Frey B, Busse MD, Schütz M, Risch AC. 2011. Effects of wild boar (Sus scrofa L.) rooting on the bacterial community structure in mixed-hardwood forest soils in Switzerland. Eur J Soil Biol 47:296–302. http://dx.doi.org/10.1016/j.ejsobi.2011.07.003
- Zumsteg A, Luster J, Göransson H, Smittenberg RH, Brunner I, Bernasconi SM, Zeyer J, Frey B. 2012. Bacterial, Archaeal and Fungal Succession in the Forefield of a Receding Glacier. Microb Ecol 63:552–64. https://doi.org/10.1007/s00248-011-9991-8

ecosystem-coupling

display_name: ecosystem-coupling

ecosystem-functions

display_name: ecosystem-functions

ecosystem-multifunctionality

display_name: ecosystem-multifunctionality

exclusion

display_name: exclusion

grassland

display_name: grassland

grazing

display_name: grazing

invertebrates

display_name: invertebrates

subalpine

display_name: subalpine

swiss-national-park

display_name: swiss-national-park

vertebrates

display_name: vertebrates

educ

en: Education, culture and sport

envidat

en: EnviDat: the Environmental Data Portal

en: EnviDat is the WSL data portal providing unified and managed access to environmental monitoring and research data. The portal has the capability to host and publish data sets. Through the sharing of unique and high-value data sets worldwide, EnviDat fosters research and contributes to advancing environmental science.

en: Ecosystem coupling and multifunctionality - exclosure experiment


maintainer: Michael Eisenring

en: european-beech alan light-pollution phytochemistry insect-herbivores

state: active

type: dataset

en: Raw_data_Ceppi et al..xlsx

en: Raw data for the Publication Ceppi et al. 2025 in Journal of Applied Ecology: ARTIFICAL NIGHT LIGHT INTENSITY MODULATES HERBIVORY AND PHYTOCHEMISTRY IN EUROPEAN BEECH

format: XLS

en: Raw data on plant morphological traits, phytochemistry and herbivory that were analyzed in the study entitled: Artificial night light intensity modulates herbivory and phytochemistry in European beech

alan

display_name: alan

european-beech

display_name: european-beech

insect-herbivores

display_name: insect-herbivores

light-pollution

display_name: light-pollution

phytochemistry

display_name: phytochemistry

educ

fr: Éducation, culture et sport

de: Bildung, Kultur und Sport

it: Istruzione, cultura e sport

fr: EnviDat: le portail de données environnementales

de: EnviDat: das Umwelt-Datenportal

it: EnviDat: il portale dei dati ambientali

fr: EnviDat est le portail de données du WSL; il offre un accès uniformisé et réglementé aux données de surveillance et de recherche en matière d’environnement. Ce portail est capable d’héberger et de publier des ensembles de données précieuses et uniques, qui sont partagées aux quatre coins de la planète. Ce faisant, EnviDat favorise la recherche et contribue à l’avancement des sciences environnementales.

de: EnviDat ist das Datenportal der WSL für einen einheitlichen und verwalteten Zugang zu Umweltdaten aus Monitoring- und Forschungsprojekten. Auf dem Portal können Datensätze gehostet und veröffentlicht werden. Durch die weltweite Bereitstellung einzigartiger und wertvoller Datensätze leistet EnviDat einen Beitrag zur Forschung im Bereich der Umweltwissenschaften.

it: EnviDat è il portale dati del WSL che fornisce un accesso unificato e gestito ai dati di monitoraggio ambientale e di ricerca. Il portale ha la capacità di ospitare e pubblicare serie di dati preziosi e unici che vengono condivisi in tutto il mondo. Così facendo, EnviDat promuove la ricerca e contribuisce al progresso della scienza ambientale.

political_level: other

wsl

capacity: public

type: organization

approval_status: approved

key: guid

key: uri

value: Envidat Harvester

en: Raw data - Artificial night light intensity modulates herbivory and phytochemistry in European beech


maintainer: Dirk Nikolaus Karger

en: precipitation climatology bioclim free-climate-data climate climate-layers climate-change temperature

en: CHELSA V1.2 (deprecated)

en: CHELSA Version 1.2 based on ERA-Interim forcings and CMIP5

format: GeoTIFF

en: CHELSA Technical Documentation

media_type: application/pdf

en: Technical documentation document including the variable names, units, scaling, offset, and background.

format: PDF

en: CHELSA V2.1 (current)

en: CHELSA Version 2.1 based on ERA5 forcings and CMIP6

en:
High-resolution information on climatic conditions is essential to many applications in environmental and ecological sciences. Here we present the CHELSA (Climatologies at high resolution for the earth’s land surface areas) data of downscaled temperature and precipitation to a high resolution of 30 arc sec. The temperature algorithm is based on statistical downscaling of atmospheric temperatures. The precipitation algorithm incorporates orographic predictors including wind fields, valley exposition, and boundary layer height, with a subsequent bias correction.

CHELSA data published in EnviDat includes the deprecated version 1.2 (originally published under 10.5061/dryad.kd1d4). Please use the current 2.1 version.

Paper Citation:
 > _Karger DN. et al. Climatologies at high resolution for the earth’s land surface areas, Scientific Data, 4, 170122 (2017) [doi: 10.1038/sdata.2017.122](https://doi.org/10.1038/sdata.2017.122)._

bioclim

display_name: bioclim

climate

display_name: climate

climate-change

display_name: climate-change

climate-layers

display_name: climate-layers

climatology

display_name: climatology

free-climate-data

display_name: free-climate-data

precipitation

display_name: precipitation

temperature

display_name: temperature


maintainer: Christian Rixen

en: stillberg treeline dwarf-shrubs tree-rings fertilization nitrogen-isotope

en: Metadata Nutrient Addition Stillberg Experimental Design

media_type: image/png

en: Nutrient addition map of the study area with the investigated plots. Each circle (study block) contains one control plot and two plots that received NPK fertilizer corresponding to 15 (NPK15) or 30 (NPK30) kg N ha-1 a-1. Contour lines specify the elevation (m a.s.l.) in steps of 10 m.

format: PNG

en: Data Nutrient Addition Stillberg

en: This link leads to the figshare repository, which contains all nutrient addition data.

format: SERVICE

en: Project Description Nutrient Addition Stillberg

media_type: text/html

en: Link to Nutrient Addition project description on the WSL website (in German).

format: HTML

en:
 Background information

The availability of nitrogen (N) and phosphorus (P) is considered to be a major factor limiting growth and productivity in terrestrial ecosystems globally. This project aimed to determine whether the growth stimulation documented in previous short‐term fertilisation trials persisted in a longer‐term study (12 years) in the treeline ecotone, and whether possible negative effects of nutrient addition offset the benefits of any growth stimulation.
Over the course of the 12 study years, NPK fertiliser corresponding to 15 or 30 kg N ha−1 a−1 was added annually to plots containing 30‐year‐old *Larix decidua* or 32‐year-old *Pinus uncinata* individuals with an understorey of mainly ericaceous dwarf shrubs. To quantify growth, annual shoot increments of trees and dwarf shrubs as well as radial growth increments of trees were measured. Nutrient concentrations in the soil were also measured and the foliar nutritional status of trees and dwarf shrubs was assessed.

 Experimental design

Over an elevation gradient of 140 m across the treeline afforestation site Stillberg, 22 locations were chosen that covered the whole range of microenvironmental conditions (*see* Nutrient addition experimental design.png). Half of the blocks included European larch (*L. decidua*) and the other half included mountain pine (*P. uncinata*). Within each block, three plantation quadrats were randomly selected as experimental plots and each plot was assigned to a control (no fertilisation) or to one of two fertiliser dose treatments (15 kg and 30 kg N ha−1 a−1). Treatments were assigned randomly but confined so that the location of fertilised plots within a block was not directly above control plots to avoid nutrient input from drainage. For details about the experiment, *see* Möhl et al (2019).

 Data description

The available datasets contain climate variables (2004-2016), nutrient isotope measurements (2010 & 2016), shrub growth measurements (2004-2016), soil parameter measurements and annual ring and shoot measurements (2004-2016). All data can be found here:
<https://figshare.com/articles/dataset/Twelve_years_of_low_nutrient_input_stimulates_growth_of_trees_and_dwarf_shrubs_in_the_treeline_ecotone/7025858>

dwarf-shrubs

display_name: dwarf-shrubs

fertilization

display_name: fertilization

nitrogen-isotope

display_name: nitrogen-isotope

stillberg

display_name: stillberg

tree-rings

display_name: tree-rings

treeline

display_name: treeline

educ

en: Education, culture and sport

en: Nutrient addition experiment at the Alpine treeline site Stillberg, Switzerland


maintainer: Avalanche Warning Service SLF

en: Avalanche fatalities per year in Switzerland since 1936/1937

media_type: text/csv

format: CSV

mimetype: text/csv

en:
Attention: this data is not updated after 2022 anymore.
This dataset contains the statistics on the number of avalanche fatalities per hydrological year in Switzerland. The data set commences with the beginning of the hydrological year 1936/37 on 01/10/1936. After the completion of a hydrological year, the new data is appended to the existing dataset.
The following information is contained (by column and column title):
 - hydrological year
 - number of fatalities in the backcountry (=tour)
 - number of fatalities in terrain close to ski areas (=offpiste)
 - number of fatalities on transportation corridors including ski runs, roads, railway lines (=transportation.corridors)
 - number of fatalities in or around buildings or in settlements (= buildings)
 - sum (of all four categories)
The definition for these four categories as described in the guidelines to the avalanche accident database:
**tour**:
activities include back-country ski, snowboard or snow-shoe touring
**offpiste**:
access from ski area, generally from the top of a skilift with short hiking distances
**transportation.corridors** ([Techel et al., 2016](http://www.geogr-helv.net/71/147/2016/ )):
people travelling or recreating on open or temporarily closed
transportation corridors (e.g. a road user or a skier on a ski run) and people working on open or closed transportation corridors (e.g. maintenance crews on roads, professional rescue teams)
**buildings** ([Techel et al., 2016](http://www.geogr-helv.net/71/147/2016/ )):
people inside or just outside buildings, and workers on high alpine building
sites

avalanche-accident-statistics

display_name: avalanche-accident-statistics

avalanche-accidents

display_name: avalanche-accidents

avalanche-fatalities

display_name: avalanche-fatalities

fatalities-in-or-around-buildings

display_name: fatalities-in-or-around-buildings

fatalities-in-settlements

display_name: fatalities-in-settlements

fatalities-in-the-backcountry

display_name: fatalities-in-the-backcountry

fatalities-on-transportation-corridors

display_name: fatalities-on-transportation-corridors

envidat

en: EnviDat: the Environmental Data Portal

en: EnviDat is the WSL data portal providing unified and managed access to environmental monitoring and research data. The portal has the capability to host and publish data sets. Through the sharing of unique and high-value data sets worldwide, EnviDat fosters research and contributes to advancing environmental science.

en: Number of avalanche fatalities per hydrological year in Switzerland since 1936-1937


maintainer: Joan Casanelles Abella

en: antwerp osmia-bicornis tartu dna-metabarcoding zurich urban-ecology osmia-cornuta poznan paris hylaeus-communis cavity-nesting-bees plant-pollinator-interactions chelostoma-florisomne trap-nest

en: Information on the study sites

en: For each site, IDs, city, coordinates, structure where trapnest was attahced and height of the trapnest is provided

en: Larval food composition

en: Explanation data variables

en: Urbanization poses threats and opportunities for the biodiversity of wild bees. A main gap relates to the food preferences of wild bees in urban ecosystems, which usually harbour large numbers of plant species, particularly at the larval stage. This data sets describes the larval food of four wild bee species (i.e. Chelostoma florisomne, Hylaeus communis, Osmica bicornis and Osmia cornuta) and three genera (i.e. Chelostoma sp., Hylaeus sp, and Osmia sp.) common in urban areas in five different European cities (i.e. Antwerp, Paris, Poznan, Tartu and Zurich). This data results from a European-level study aimed at understanding the effects of urbanization on biodiversity across different cities and citiscapes, and a Swiss project aimed at understanding the effects of urban ecosystems in wild bee feeding behaviour. Wild bees were sampled using standardized trap-nests in 80 sites (32 in Zurich and 12 in each of the remaining cities), selected following a double gradient of available habitat at local and landscape scales. Larval pollen was obtained from the bee nests and identified using DNA metabarconding. The data provides the plant composition at the species or genus level of the different bee nests of the studied species in the studied sites of the five European cities. For Hylaeus communis, this is the first study in reporting larval food composition.

antwerp

display_name: antwerp

cavity-nesting-bees

display_name: cavity-nesting-bees

chelostoma-florisomne

display_name: chelostoma-florisomne

dna-metabarcoding

display_name: dna-metabarcoding

hylaeus-communis

display_name: hylaeus-communis

osmia-bicornis

display_name: osmia-bicornis

osmia-cornuta

display_name: osmia-cornuta

paris

display_name: paris

plant-pollinator-interactions

display_name: plant-pollinator-interactions

poznan

display_name: poznan

tartu

display_name: tartu

trap-nest

display_name: trap-nest

urban-ecology

display_name: urban-ecology

zurich

display_name: zurich

en: Larval food composition of four wild bee species in five European cities


en: periurban urban-biodiversity pest-invasion biodiversity urban-ecology biodiversity-experiment birds insect-herbivores urban

en:
We established our study sites in the cities of Basel, Lugano and Zurich, and their peri-urban forests. We characterised urban tree cover of each city using a rectangular grid with squares of 100x100 m. Within each square, we measured the area covered by urban trees using the European Union's Copernicus Land Monitoring Service information, Urban Atlas Street Tree Layer 2018 [https://doi.org/10.2909/205691b3-7ae9-41dd-abf1-1fbf60d72c8c](https://doi.org/10.2909/205691b3-7ae9-41dd-abf1-1fbf60d72c8c).
Then we assigned each square to four categories of urban tree cover that roughly represented the main types of urban uses:
1) low cover, industrial/commercial areas, 0-20% tree cover;
2) intermediate cover, residential areas, 20-40% tree cover;
3) high cover, urban parks and cemeteries, 40-60% tree cover;
4) peri-urban forests (natural/semi-natural forests), 60-100% tree cover.

Our study sites consisted of eight 1.2 km-long transects in each city, equally distributed across the four tree cover categories. We designed an experimental measurements of bird predation rate on herbivore insect larvae using artificial caterpillars. They mimicked a variety of common, cryptically-coloured herbivorous species, which are usually preyed by birds. We designed caterpillars with green water-resistant modelling clay (Staedtler Noris®), 4-6 cm long and approximately 0.5 cm wide. Caterpillars were threaded to a thin (Ø = 0.6 mm) zinc-plated iron wire and tethered to branches of trees or bushes at a height up to 2 m from the ground. We placed 2-4 caterpillar mimics every 200 m along each transect (five points per transect) or, when not possible due to absence of suitable branches, within 200 m from the point.
In total, we placed 270 caterpillar mimics at 115 points during the late breeding season (June-July 2023). Caterpillar mimics were checked after 10-14 days and replaced with new ones at new locations within 10 m from the previous ones. During each of the two visits, we counted the number of bird predation marks.

biodiversity

display_name: biodiversity

biodiversity-experiment

display_name: biodiversity-experiment

birds

display_name: birds

insect-herbivores

display_name: insect-herbivores

periurban

display_name: periurban

pest-invasion

display_name: pest-invasion

urban

display_name: urban

urban-biodiversity

display_name: urban-biodiversity

en: Urban bird predation on artificial caterpillars


maintainer: Elisabeth D. Hafner

en: avalanche-activity avalanche remote-sensing aerial-imagery avalanche-mapping

en: 1999 avalanche data

media_type: application/zip

en:
**for English see below**
Dieser Datensatz enthält die **Umrisse der 11'120 Lawinen** die aus schwarzweiss Luftbildern, welche zwischen dem 25.2.1999 und dem 1.3.1999 aufgenommen wurden, kartiert wurden. Die Lawinenumrisse haben verschiedene Attribute welche im **beigelegten Beispielschlüssel** beschrieben sind (Beispielschluessel_1999_d.pdf).
Es gibt drei Shapefiles:
* **avalanches1999_endversion1.shp**: die kartierten Lawinen (für Attribute siehe Beispielschlüssel!)
* **area_images_1999.shp**: Fläche die durch die Luftbilder abgedeckt wurde
* **clouds_1999.shp**: grobe Umrisse der Wolken in den Bildern
#
 *******************************************************************************
This dataset contains the **outlines of 11'120 avalanches** mapped from panchromatic aerial imagery taken between February 25, 1999 and March 1, 1999. The avalanche outlines have various attributes which are described in the **attached example key** (ExampleKey_AvalMapping_1999_e.pdf).
There are three shapefiles:
* **avalanches1999_endversion1.shp**: the mapped avalanches (for attributes see example key!)
**area_images_1999.shp**: Area covered by the aerial images
**clouds_1999.shp**: rough outlines of the clouds in the images

format: ZIP

en:
**for English see below**
*******************************************************************************

 Datenbeschrieb

Dieser Datensatz enthält die **Umrisse der 11'120 Lawinen** die aus schwarzweiss Luftbildern, welche zwischen dem 25.2.1999 und dem 1.3.1999 aufgenommen wurden, kartiert wurden. Die Lawinenumrisse haben verschiedene Attribute welche im **beigelegten Beispielschlüssel** beschrieben sind (Beispielschluessel_1999_d.pdf).

Es gibt drei Shapefiles:
* **avalanches1999_endversion1.shp**: die kartierten Lawinen (für Attribute siehe Beispielschlüssel!)
* **area_images_1999.shp**: Fläche die durch die Luftbilder abgedeckt wurde
* **clouds_1999.shp**: grobe Umrisse der Wolken in den Bildern

 Bildverfügbarkeit:

Die entzerrten Luftbilder können von der [Swisstopo](https://s.geo.admin.ch/rpq3hufnb588) einzeln über den Downloadlink heruntergeladen werden (herzlichen Dank an Holger Heisig für die Prozessierung und ans BAFU für die Finanzierung der Prozessierung!!). Über "Erweiterte Werkzeuge"/ "Datei importieren" können die Bilder nach hereinziehen des Orthofotolinks direkt im map.geo.admin.ch angezeigt werden.

 Weiterführende Litertaur:
Details zu den Luftbildern und zur Kartierung, sowie eine kleine Analyse und Empfehlungen zur Verwendung der Daten finden sich unter

- **FAN Publikation** (Info folgt)

- Dal, F. J., Hafner, E. D., Peters, T., Narnhofer, D., Caye Daudt, R., Heisig, H., & Bühler, Y. (2024). Automated snow avalanche mapping with deep learning in aerial imagery from the extreme avalanche winter of 1999. In K. Gisnås, P. Gauer, H. Dahle, M. Eckerstorfer, A. Mannberg, & K. Müller (Eds.), Proceedings of the international Snow Science Workshop 2024 (pp. 1264-1271). Norwegian Geotechnical Institute.

- Hafner, E. D., Techel, F., Heisig, H., Dal, J. F., & Bühler, Y. (2024). Remotely sensed avalanche activity during three extreme avalanche periods in Switzerland. In K. Gisnås, P. Gauer, H. Dahle, M. Eckerstorfer, A. Mannberg, & K. Müller (Eds.), Proceedings of the international Snow Science Workshop 2024 (pp. 1222-1229). Norwegian Geotechnical Institute.

*******************************************************************************

 Data description

This dataset contains the **outlines of 11'120 avalanches** mapped from panchromatic aerial imagery taken between February 25, 1999 and March 1, 1999. The avalanche outlines have various attributes which are described in the **attached example key** (ExampleKey_AvalMapping_1999_e.pdf).

There are three shapefiles:
* **avalanches1999_endversion1.shp**: the mapped avalanches (for attributes see example key!)
**area_images_1999.shp**: Area covered by the aerial images
**clouds_1999.shp**: rough outlines of the clouds in the images

 Image availability:
The rectified aerial images can be downloaded individually from [Swisstopo](https://s.geo.admin.ch/bg6beetinfdc) via the download link (many thanks to Holger Heisig for processing and to BAFU for financing the processing!!). The images can be displayed directly in map.geo.admin.ch via “Advanced tools”/“Import file” after dragging in the orthophoto link.

 Further Reading:
Details on the aerial images and mapping, as well as a brief analysis and tips on using the data can be found at

- **FAN Publication** (info to follow)

- Dal, F. J., Hafner, E. D., Peters, T., Narnhofer, D., Caye Daudt, R., Heisig, H., & Bühler, Y. (2024). Automated snow avalanche mapping with deep learning in aerial imagery from the extreme avalanche winter of 1999. In K. Gisnås, P. Gauer, H. Dahle, M. Eckerstorfer, A. Mannberg, & K. Müller (Eds.), Proceedings of the international Snow Science Workshop 2024 (pp. 1264-1271). Norwegian Geotechnical Institute.

- Hafner, E. D., Techel, F., Heisig, H., Dal, J. F., & Bühler, Y. (2024). Remotely sensed avalanche activity during three extreme avalanche periods in Switzerland. In K. Gisnås, P. Gauer, H. Dahle, M. Eckerstorfer, A. Mannberg, & K. Müller (Eds.), Proceedings of the international Snow Science Workshop 2024 (pp. 1222-1229). Norwegian Geotechnical Institute.

aerial-imagery

display_name: aerial-imagery

avalanche

display_name: avalanche

avalanche-activity

display_name: avalanche-activity

avalanche-mapping

display_name: avalanche-mapping

remote-sensing

display_name: remote-sensing

educ

en: Education, culture and sport

en: Avalanche outlines February and March 1999 from aerial imagery


photogrammetric-drone-data-gruenbodeli

maintainer: EnviDat Support

title_for_slug: photogrammetric-drone-data-gruenbodeli

en: orthophoto snow-depth-mapping wingtra dsm photogrammetry uav drone-data

en: Data Access

en:
- ortho (3 cm/25 mm resolution)
- DSM (10 cm resolution)
- HS (10 cm resolution)
- RGB images
- Agisoft Metashape Project Report

All products are also available as a cloud optimized geotiff.

en:
We conducted various drone flights at Grüenbödeli near Davos with the Sony RX1R II mounted on a Wingtra drone during 2020/21/22. The data was processed with the Agisoft Metashape Professional Software.
The following products are available for download:
- DSM 10cm resolution
- Orthomosaic 3cm/25mm resolution
- Snow Raster 10cm resolution
- original RGB images

drone-data

display_name: drone-data

dsm

display_name: dsm

orthophoto

display_name: orthophoto

photogrammetry

display_name: photogrammetry

snow-depth-mapping

display_name: snow-depth-mapping

uav

display_name: uav

wingtra

display_name: wingtra

envidat

en: EnviDat: the Environmental Data Portal

en: EnviDat is the WSL data portal providing unified and managed access to environmental monitoring and research data. The portal has the capability to host and publish data sets. Through the sharing of unique and high-value data sets worldwide, EnviDat fosters research and contributes to advancing environmental science.

en: Photogrammetric Drone Data Grüenbödeli


en: water-isotopes forest critical-zone cryogenic-extraction hydrogen-isotopes ecohydrology europe isotopes picea-abies hydrology fagus-sylvatica soil-water spruce oxygen-isotopes beech xylem-water

en: Dataset containing the metadata of the sampling sites, including site-, soil-, tree-specific information.

en: Dataset containing information about sample weights, cryogenic water extraction and the actual hydrogen and oxygen isotope data of the study.

en: Dataset containing the information on the canopy cover.

en: WATSON_Canopy_Pictures (individual JPG files)

en: All canopy pictures as individual files.

en: All canopy pictures in one ZIP file.

en: Here, we present a comprehensive isotope dataset of soil and stem xylem water collected during two pan-European sampling campaigns in spring and summer 2023. The dataset includes samples from 40 forest sites featuring beech (Fagus sylvatica), spruce (Picea abies), or mixed beech-spruce forests, and is supplemented with detailed site-specific, soil-specific, and tree-specific metadata. Samples and metadata were collected by various researchers across Europe following a standardized protocol. Soil samples were taken at up to five depths (ranging from 0 to 90 cm), and stem xylem samples were collected from three beech and/or spruce trees at each site. All samples were sent to a single laboratory for analysis, where water was extracted using cryogenic vacuum distillation and analyzed with an isotope laser spectrometer. Additionally, a subset of the samples underwent analysis with an isotope ratio mass spectrometer. The complete description of the data can be found in a publication in Earth System Science Data (in review).

beech

display_name: beech

critical-zone

display_name: critical-zone

cryogenic-extraction

display_name: cryogenic-extraction

ecohydrology

display_name: ecohydrology

europe

display_name: europe

fagus-sylvatica

display_name: fagus-sylvatica

forest

display_name: forest

hydrogen-isotopes

display_name: hydrogen-isotopes

hydrology

display_name: hydrology

isotopes

display_name: isotopes

oxygen-isotopes

display_name: oxygen-isotopes

picea-abies

display_name: picea-abies

soil-water

display_name: soil-water

spruce

display_name: spruce

water-isotopes

display_name: water-isotopes

xylem-water

display_name: xylem-water

en: Soil and stem xylem water isotope data from two pan-European sampling campaigns


maintainer: Stephan Zimmermann

en: calcium biogeochemical-cycling fagus-orientalis-lipsky magnesium potassium

en: Concentrations of Ca, Mg, K, and Na in throughfall, litterflow and soil solution of a Hyrcanian forst in Northern Iran

en:
Throughfall, litterflow and soil solution were sampled during one whole year under five Oriental beech trees in a mixed Hyrcanian beech forest. The amounts of Ca2+, Mg2+, K+ and Na+ in these fluxes were calculated based on their concentrations and the sampled volumes, and subsequently compared with the respective fluxes in the rainfall and soil solution of an adjacent forest gap.
In addition six soil profiles, one close to every single tree and one in the forest gap, were analyzed for pH, CaCO3, organic matter and texture.

biogeochemical-cycling

display_name: biogeochemical-cycling

calcium

display_name: calcium

fagus-orientalis-lipsky

display_name: fagus-orientalis-lipsky

magnesium

display_name: magnesium

potassium

display_name: potassium

educ

en: Education, culture and sport

en: Base cation dynamics in an Oriental beech forest


en: urban urban-biodiversity fungi urban-ecology biodiversity birds insects fungal-pathogen

en: Data tables including individual bird data with biometry, prey species from faecal samples, and fungi species from swab samples.

en:
We captured birds in urban parks and peri-urban forests in the cities of Lugano and Zurich. The urban sites were located in an urban forest (Lugano) and in a cemetery (Zurich), and the bird assemblages were comparable with those of the surrounding forests.
We captured individual birds during sessions lasting multiple days using mist-nets, concentrating the activities within 3-4 hours after sunrise and before sunset. We extracted birds from nets using single-use nitrile gloves and gently wiped their body parts with a dry foam swab MW940 (Medical Wire & Equipment), which was then placed in a screw-capped tube (2 ml), which was previously filled with 1 ml ethanol puriss. (Sigma-Aldrich) under sterile conditions.Then, we marked each bird with an alphanumeric leg band, identified the species, determined the sex and age whenever possible, measured for relevant biometrics (e.g. wing chord), the amount of body fat and pectoral muscle (i.e. body condition), and weighed.
Afterwards, we placed each bird in a disposable paper bag within a cotton bag that ensured respiration and left it for few minutes in a safe and calm place to allow for defecation and the collection of faecal samples. The faecal sample was collected from insectivorous and omnivorous birds, and few granivorous birds that are known to occasionally eat invertebrates, such as the house sparrow Passer domesticus.
After releasing the bird, we stored the faecal sample in a screw-capped tube (2 ml), which was previously filled with 1 ml ethanol puriss. (Sigma-Aldrich) under sterile conditions. We wore single-use nitrile gloves and used sterile toothpicks to collect solid samples. The tubes were initially stored in the refrigerator during the field campaign and no later than seven days after collection at -20°C until prepared for DNA extraction.
DNA extraction and library preparation was performed at WSL Phytopathology facilites, Sequencing was performed at Lausanne Genomic Technologies Facility (University of Lausanne; [https://wp.unil.ch/gtf/](https://wp.unil.ch/gtf/)) on an AVITI benchtop sequencing system (Element Biosciences). Bioinformatics was performed at the Genetic Diversity Center (ETH Zurich) using a custom pipeline.

biodiversity

display_name: biodiversity

birds

display_name: birds

fungal-pathogen

display_name: fungal-pathogen

fungi

display_name: fungi

insects

display_name: insects

urban

display_name: urban

urban-biodiversity

display_name: urban-biodiversity

urban-ecology

display_name: urban-ecology

envidat

en: EnviDat: the Environmental Data Portal

en: Urban bird capture data with prey species and fungal community


weissfluhjoch-dataset-for-esm-snowmip

maintainer: Nander Wever

title_for_slug: weissfluhjoch-dataset-for-esm-snowmip

en: snow-depth weissfluhjoch snow-models esm-snowmip snow

en: ini file

en: This ini file is required by data-converter from MeteoIO version 1887 (http://models.slf.ch/p/meteoio/)

mimetype: application/zip

en: Conversion script

en:
Use this script to convert WFJ_optimaldataset_v6.smet to wfj.smet, the file that is used for the ESM-SnowMIP project. It also requires the file: WFJ_AirPressure_Data.zip

Use: bash create_wfj_dataset.sh > input/wfj.smet

mimetype: text/x-sh

en: Dataset Weissfluhjoch for ESM-SnowMIP

en: This Weissfluhjoch dataset is a processed version of the Weissfluhjoch dataset version 6 from https://doi.org/10.16904/6. This dataset was specially created for the ESM-SnowMIP project. Here it is documented how this dataset has been created.

esm-snowmip

display_name: esm-snowmip

snow

display_name: snow

snow-depth

display_name: snow-depth

snow-models

display_name: snow-models

weissfluhjoch

display_name: weissfluhjoch

en: EnviDat is the WSL data portal providing unified and managed access to environmental monitoring and research data. The portal has the capability to host and publish data sets. Through the sharing of unique and high-value data sets worldwide, EnviDat fosters research and contributes to advancing environmental science.

en: Weissfluhjoch dataset for ESM-SnowMIP


defoliation-growth-data

title_for_slug: defoliation-growth-data

en: literature-review tree-growth defoliation relative-growth forest-health

en:
Using a literature review extracted data from 26 published papers which had data on individual tree defoliation and growth relationships In Europe.
This data set includes 415 defoliation-growth data points from 16 different tree species covering 15+ countries in Europe. Growth metrics used in individual papers were different, and included increments in terms of volume, tree rings width, radial growth, basal area and biomass. All growth metrics were converted into relative growth (presented in this dataset), expressed as a percentage of the growth of progressively defoliated trees in relation to the growth of those trees adopted as reference (control) defoliation level (i.e. the growth of the lowest defoliation level in each study was used as the reference growth point).

defoliation

display_name: defoliation

forest-health

display_name: forest-health

literature-review

display_name: literature-review

relative-growth

display_name: relative-growth

tree-growth

display_name: tree-growth

en: Defoliation-Growth data


en: snowmelt imis monitoring phenology alpine long-term grassland ultrasonic snow climate

en: Contains custom functions required for the scripts; this file must be sourced before running other scripts.

format: OCTET

en: Contains unfiltered annual phenological, temperature and snow-climatic data for all IMIS stations.

en: Contains filtered annual phenological data from 40 IMIS stations suitable and used for phenological analysis

en: Is the readme that documents all data and code in this repository

en: Snow3.png and grass.png are needed for plotting the methods plot in 5_methods_plots.Rmd

en: Documents and reproduces the statistical models used in the analysis.

en: implements the phenofit method. Note: This script is computationally intensive and may take a while to run. We recommend using the temporary data created from it (i.e. starting from 2_temperature.R). For a better description of the phenofit R package and the different methods to determine the SOG, see Kong et al. (2022): https://doi.org/10.1111/2041-210X.13870

en: Generates the methods-related figures used in the publication.

en: Temporary data. Contains annual unfiltered output of phenofit (i.e. the variable names are in line with the phenofit documentation), including all curve parametrization variables for plotting and additional variables, which are not necessarily used in the analysis.

en: Contains IMIS station location data

en: Contains daily climate station data including daily median temperatures, snow depth, plant height, and annual melt-out timings.

en: Contains annual phenological variables and phenofit outputs, sorted, ordered and renamed according with the readme in this repository

en:
Long-term phenological data in alpine regions are often limited to a few locations and thus, little is known about climate-change induced plant phenological shifts above the treeline. Because plant growth initiation in seasonally snow-covered regions is largely driven by snowmelt timing and local temperature, it is essential to simultaneously track phenological shifts, snowmelt, and near-ground temperatures. In this study, we make use of ultrasonic snow height sensors installed at climate stations in the Swiss Alps to reveal phenological advance of grassland ecosystems and relate them to climatic changes over 25 years (1998 – 2023). When snow is absent, these snow height sensors additionally provide information on plant growth at a uniquely fine temporal scale. Daily temperatures and snow- / plant height values, annual phenological- and melt-out dates were derived from IMIS climate station data (doi.org/10.16904/envidat.406). We observed an advance of green-up by -2.4 days/decade coinciding with strong warming of up to +0.8°C/decade. Although the timing of melt-out has not changed significantly over the study period in this focal region, phenological responses to early melt-out years varied due to differing influences of photoperiodic and thermal constraints, which were not equally important across elevations and communities. Phenological shifts of alpine grasslands are thus likely to become even more pronounced if melt-out timing advances in the future as predicted.
This repository relates to the publication Zehnder et al. (2025). Snow height sensors reveal phenological advance in alpine grasslands. Global Change Biology. accepted. The model data and code that support our findings are deposited in this repository.

alpine

display_name: alpine

climate

display_name: climate

grassland

display_name: grassland

imis

display_name: imis

long-term

display_name: long-term

monitoring

display_name: monitoring

phenology

display_name: phenology

snowmelt

display_name: snowmelt

ultrasonic

display_name: ultrasonic

educ

en: Education, culture and sport

en: IMIS Phenology Data Since 1998


urban-birds-point-counts-data

title_for_slug: urban-birds-point-counts-data

en: environmental-gradients birds spatially-distributed-sampling urban periurban biodiversity

en:
The study area included the cities of Basel, Lugano and Zurich. The sampling design reflected four main city districts related to four categories of tree cover (Figure 1): 1) industrial/commercial areas, 0-20% tree cover; 2) residential areas, 20-40% tree cover; 3) urban parks and cemeteries, 40-60% tree cover; 4) peri-urban forests (natural/semi-natural forests), 60-100% tree cover. The tree cover gradient was evaluated across a square grid with a square side length of 100 m. For each square, we evaluated the area covered by urban trees using the Urban Atlas Street Tree Layer 2018 (European Union’s Copernicus Land Monitoring Service information, 2021).
In each city, we established eight transects of 1.2 km, two for each tree cover category. Transects crossed contiguous grid squares belonging to the same tree cover categories as much as possible. We performed aural-visual point counts of birds every 200 m along each transect (six points per transect) during the breeding season (April-June 2024) (Bibby et al., 2000). The transects were walked at the time of highest activity of birds, which is from thirty minutes after sunrise until 4 hours later. Each transect was walked twice, 20-60 days apart, to account for imperfect detection of birds and sample completeness (Balestrieri et al., 2017; Kéry et al., 2005).

biodiversity

display_name: biodiversity

birds

display_name: birds

environmental-gradients

display_name: environmental-gradients

periurban

display_name: periurban

spatially-distributed-sampling

display_name: spatially-distributed-sampling

urban

display_name: urban

envidat

en: EnviDat: the Environmental Data Portal

en: EnviDat is the WSL data portal providing unified and managed access to environmental monitoring and research data. The portal has the capability to host and publish data sets. Through the sharing of unique and high-value data sets worldwide, EnviDat fosters research and contributes to advancing environmental science.

en: Urban birds point counts data


maintainer: Yohann Chauvier

en: taxonomy range-maps filtering synonyms flags occurrence-records landsca gbif r-package species-observations conservation ecoregions large-scale species-distribution accepted-name hard-limit

en: gbif.range R package version 1.1 (current)

en: wsl.gbif R package version 0.1 (deprecated)

en: Test version, not to be used.

en: GitHub URL

en: gbif.range R package version 0.2 (deprecated)

en: Although species range may be obtained using expert maps or modeling methods, expert data is often species-limited and statistical models need more technical expertise as well as many species observations. When unavailable, such information may be extracted from the Global Biodiversity Information facility (GBIF), the largest public data repository inventorying georeferenced species observations worldwide. However, retrieving GBIF records at large scale may be tedious if users are unaware of specific tools and functions that need to be employed. Here we present *gbif.range*, an R library that contains automated methods to generate species range maps from scratch using in-house ecoregions shapefiles and an easy-to-use GBIF download wrapper. Finally, this library also offers a set of additional very useful parameters and functions for large GBIF datasets (generate doi, extract GBIF taxonomy, records filtering...). [gbif.range R project](https://github.com/8Ginette8/gbif.range)

accepted-name

display_name: accepted-name

conservation

display_name: conservation

ecoregions

display_name: ecoregions

filtering

display_name: filtering

flags

display_name: flags

gbif

display_name: gbif

hard-limit

display_name: hard-limit

landsca

display_name: landsca

large-scale

display_name: large-scale

occurrence-records

display_name: occurrence-records

r-package

display_name: r-package

range-maps

display_name: range-maps

species-distribution

display_name: species-distribution

species-observations

display_name: species-observations

synonyms

display_name: synonyms

taxonomy

display_name: taxonomy

en: gbif.range - An R package to generate species range maps based on ecoregions and a user-friendly GBIF wrapper


maintainer: Marco Moretti

en: beekeeping resource-availability urban-biodiversity urbanization honeybees pollinator-experiment city4bees wild-bees bettergardens zurich gardens urban-ecosystems

en: Metadata for the predictor and response dataset

en: Response and predictors used for the SEMs

en: Metadata for the predictors and response dataset

en:
Mean, standard deviation , minimum and maximum values of six functional traits. The data has the following structure:
- Bee_species: bee scientific name
- mean_trait: mean value
- sd_trait: standard deviation
_min_trait: minimum value
- max_trait: maximum value
- trait: the six traits used. ITD = Intertegular distance (mm); Pheno.duration = duration of the phenology; Pheno.start = start of the phenology; Pheno.end = end of the phenology; Rel.tongue.length= relative tongue length (mm); Daytime.activity = daily activity (h)
DistNichePart: functional dissimilarity in a multidimentional trait space defined by five traits between wild bee individuals of a wild bee species and the honeybee individuals. Values are at the species level.

en:
The data contains the following columns:

  SiteID: numeric, the identity of the garden where the bee was sampled.
  Taxonname: character, species name
  Lecty: numeric discreet, a mesure of specialisation degree. Levels: 1 = monolectic; 2=oliolectic; 3=polilectic
  Pheno.start= numeric discreet, the starting week of the flying activity.
  Pheno.end= numeric discreet, the ending week of the flying activity.
  Pheno.duration=numeric discreet, Pheno.end - Pheno.start
  Daytime.activity = numeric continuous, individual-level measure of the day hour when the bee was sampled
  ITD = numeric continuous, individual-level measure of the InterTegular Distance, a metric of mobility in mm.
  Rel.tongue.length =numeric continuous, individual-level measure of the relative tongue length (Tongue length / ITD), a metric of specilaisation degree, in mm.

en:
Cities are socio-ecological systems that filter and select species, thus establishing unique species assemblages and biotic interactions. Urban ecosystems can host richer wild bee communities than highly intensified agricultural areas, specifically in resource-rich urban green spaces such as allotment and family gardens. At the same time, urban beekeeping has boomed in many European cities, raising concerns that the fast addition of a large number of managed bees could deplete the existing floral resources, triggering competition between wild bees and honeybees. The data has been used to investigated the interplay between resource availability and the number of honeybees at local and landscape scales and how this relationship influences wild bee diversity.

This dataset contains the raw and processed data supporting the findings from the paper: "Low resource availability drives feeding niche partitioning between wild bees and honeybees in a European city".

The data contains:
1. Bee trait measurements at the species and individual-level of five functional traits.
2. The values of the feeding niche partitioning (functional dissimilarity to honeybees)
3. The predictors of resource availability and beekeeping intensity at local and landscape scales used in the modelling of the paper for the 23 experimental sites.

beekeeping

display_name: beekeeping

bettergardens

display_name: bettergardens

gardens

display_name: gardens

honeybees

display_name: honeybees

pollinator-experiment

display_name: pollinator-experiment

resource-availability

display_name: resource-availability

urban-biodiversity

display_name: urban-biodiversity

urban-ecosystems

display_name: urban-ecosystems

urbanization

display_name: urbanization

wild-bees

display_name: wild-bees

zurich

display_name: zurich

en: Niche partitioning between wild bees and honeybees


en: subalpine meteorological-data alptal first-order-catchment snow runoff

en: Daily meteorological and snow data (1968-2022) Alptal

en:
Air temperature, dew point temperature, relative humidity, wind speed, precipitation and global radiation have been measured (since 1981) on a meadow (Erlenhöhe) in the Erlenbach catchment. It is located at 1'220 m a.s.l. with an exposure towards west. In order to assess the spatial variability of precipiation, two additional rain gauges have been running in the Vogelbach and Lümpenenbach catchment, respectively.
For the period before 1981, meteorological variables have been extrapolated from the nearby MeteoSwiss station Einsiedeln (ca. 15 km north) to the WSL station Erlenhöhe using a linear correction factor for air temperature, wind speed and precipitation.
Snow depth has been recorded automatically since 2003. Snow water equivalent has been calculated using two numerical models: COUP model (Jansson, 2012) and DeltaSnow model (Winkler et al., 2021)

Jansson, P.-E. (2012). COUP model: model use, calibration, and validation.Transactions of the ASABE, 55(4), 1335–1344.
Winkler, M., et al. (2021): Snow water equivalents exclusively from snow depths and their temporal changes: the Δsnow model, Hydrol. Earth Syst. Sci., 25, 1165-1187, 10.5194/hess-25-1165-861 2021.

en: Daily runoff data from three steep torrents in the Alptal (1968-2022)

en: Runoff, water temperature and electrical conductivity have been measured since 1968 at the outlet of three torrential catchments (with an area of approximately 1 km2 each) located in the Alptal valley: Erlenbach, Lümpenenbach and Vogelbach. This data set provides daily mean values; data at 10-min resolution can be provided on request for the period 1984-2022 (contact: alptal@wsl.ch).

en: Manual snow measurements Alptal (1968-2022)

en:
Snow depth, snow density and snow water equivalent have been measured since winter 1968/69 along more than 15 transects distributed across the Alptal valley representing different altitudes, aspects and land use (closed forest and open meadows). Length of the snow courses: 20 to 30 m. The measurement interval has been weekly to monthly.
More details about the locations and methods can be found in Stähli, M.; Gustafsson, D., (2006). Long-term investigations of the snow cover in a subalpine semi-forested catchment. Hydrological Processes, 20, 411-428. doi: 10.1002/hyp.6058.

en: Snow measurements Alptal 1969-2024 EnviDat updated

en: Meteorological data Alptal 1968-2024 EnviDat updated

en:
Air temperature, dew point temperature, relative humidity, wind speed, precipitation and global radiation have been measured (since 1981) on a meadow (Erlenhöhe) in the Erlenbach catchment. It is located at 1'220 m a.s.l. with an exposure towards west. In order to assess the spatial variability of precipitation, two additional rain gauges have been running in the Vogelbach and Lümpenenbach catchment, respectively. For the period before 1981, meteorological variables have been extrapolated from the nearby MeteoSwiss station Einsiedeln (ca. 15 km north) to the WSL station Erlenhöhe using a linear correction factor for air temperature, wind speed and precipitation. Snow depth has been recorded automatically since 2003. Snow water equivalent has been calculated using two numerical models: COUP model (Jansson, 2012) and DeltaSnow model (Winkler et al., 2021)
Jansson, P.-E. (2012). COUP model: model use, calibration, and validation.Transactions of the ASABE, 55(4), 1335–1344. Winkler, M., et al. (2021): Snow water equivalents exclusively from snow depths and their temporal changes: the Δsnow model, Hydrol. Earth Syst. Sci., 25, 1165-1187, 10.5194/hess-25-1165-861 2021.

en: Runoff daily mean Alptal 1968-2024 EnviDat updated

en: Runoff, water temperature and electrical conductivity have been measured since 1968 at the outlet of three torrential catchments (with an area of approximately 1 km2 each) located in the Alptal valley: Erlenbach, Lümpenenbach and Vogelbach. This data set provides daily mean values; data at 10-min resolution can be provided on request for the period 1984-2024 (contact: alptal@wsl.ch).

en:
This data set includes 56 years of hydrometeorological measurements from small (first-order) catchments in the pre-alpine valley Alptal. Here we provide daily mean values; values in hourly or 10-min resolution can be provided on demand.
Runoff has been measured at the outlet of three small (first-order) catchments of approximately 1 km2 area: Erlenbach (two independent runoff measurements), Vogelbach and Lümpenenbach. The catchments are similar with regard to geology (Flysch) and soil conditions (clay soils), but differ in forest coverage (20 to 60%). A detailed description of the catchments can be found at https://www.wsl.ch/alptal .
Runoff in these small catchments is typically very dynamic and can temporally carry large amounts of sediment and large wood. Thus, the accuracy of the measurements at very large flow is limited.
Meteorological variables have been measured on a meadow (Erlenhöhe) located in the Erlenbach catchment at 1220 m a.s.l. using a standard meteorological station (incl. ventilated air temperature and heated rain gauges). In addition, precipitation has also been recorded at two other locations (in the Vogelbach and Lümpenenbach catchments).
Snow measurements have been conducted weekly to monthly since 1968 at more than 15 locations (30-m transects) representing different altitudes, aspects and land uses (meadow, forest). In addition, snow depth has been recorded continuously since 2003 at Erlenhöhe, and for this location we also include a simulation of snow depth and SWE (using the numerical models COUP and DeltaSnow) that assimilates the manual weekly snow-course measurements. Details on these snow measurements can be found in Stähli, M. and Gustafsson, D. 2006. Hydrol. Proc., 20, 411-428. doi: 10.1002/hyp.6058.
Further information on the methods and sensors can be found at https://www.wsl.ch/alptal .
A first version of this data set (for the period 1968-2017) was uploaded in June 2018 at the occasion of the 50-year anniversary. This original data set was updated in February 2021 (with data from 2018 and 2019), and this data set was used for a longterm trend analysis, submitted for publication in a special issue of Hydrological Processes. Further updates of the data set (with data from 2020 to 2022) were uploaded in March 2023, February 2024 and March 2025.

alptal

display_name: alptal

first-order-catchment

display_name: first-order-catchment

meteorological-data

display_name: meteorological-data

runoff

display_name: runoff

snow

display_name: snow

subalpine

display_name: subalpine

educ

en: Education, culture and sport

envidat

en: EnviDat: the Environmental Data Portal

en: EnviDat is the WSL data portal providing unified and managed access to environmental monitoring and research data. The portal has the capability to host and publish data sets. Through the sharing of unique and high-value data sets worldwide, EnviDat fosters research and contributes to advancing environmental science.

en: Longterm hydrological observatory Alptal (central Switzerland)


en: urban-ecosystems dominance anthophila cities honeybees urban-beekeeping sads wildbees

en: Dataset proportion honeybees and wild bees

en:
The dataset provide the main features for each included dataset from published and unpublished research works. Each row represents a dataset, which provides bee community data in one city / urban aggregation. Overall, it contains 68 datasets from 46 cities in 15 countries and five continents. The data is organised in the following columns:
- Article_ID: id used in the study
- DOI: doi of the publication, when available
- Authors
- Year: of publication
- Duration sampling (years)
- Start_sampling: month of the year
- End_sampling: month of the year
- Continent
- Country
- N. cities: number of cities included in the publication
- City
- Sampling type: active, passive or both
- Sampling methodology: within each sampling type.
- Habitats: list of sampling habitats
- N_habitats: number of sampling habitats
- Parks_similar: binnary (1 = yes; 0 = no)
- Agriculture: binnary (1 = yes; 0 = no)
- Private_gardens: binnary (1 = yes; 0 = no)
- Green_roofs: binnary (1 = yes; 0 = no)
- seminatural_natural_wasteland: binnary (1 = yes; 0 = no)
- other_unspecified: binnary (1 = yes; 0 = no)
- Number sites
- Abundance honeybees
- Abundance wild bees
- Total bees
- Proportion honeybees
- Proportion wildbees
- Area:. of the studied city/urban aggregation

en: R script for analyses and plots

en: Dataset species abundance

en:
This data resource provides the individual species abundance per city / urban aggregation. It contains data from 41 datasets from 37 studies collected in 33 cities / urban aggregations. It contains the following columns:
 - Article_ID
- DOI
- Authors
- Year
- Continent
- Country
- City
- Bee genus
- Bee species
- Bee merged
- Bee full
- Abundance
- Bee_ID_per_study: ID for the study

en: Dataset coordinates study cities

en:
The data contains the latitude and longitude of the studied cities / urban aggregations. It contains the following columns:
- Continent
-Country
- City
- Lat: Latitude
- Lon: Longitude

en:
The following repository contains the data and code to reproduce the figures analyses of the paper "Examining honeybee (Apis mellifera) dominance patterns within urban bee communities worldwide". It contains three datasets and one .R file

---

The data results from a literature review on abundance distribution of urban bee communities worldwide, aiming at obtaining studies reporting total abundances of honeybees and wildbees in a given city, and species abundances of wildbees and honeybees in a given city.

---

1. **Data**:
- *20240308_cities_coords.csv*: the data contains the latitude and longitude of the included cities in the study. It is used for making the world map with the cities.
- *20241201_proportion_HB_WB.csv*: the data contains the total wild bee and honeybee abundances per city/urban aggregation, and the proportions.
- *20241201_rad_bees.csv*: the data contains the species abundances per city/urban aggregation

2. **Script**:
- *HoneybeeDominance_Analyses_Figures.R*: the script to run the analyses and make the plots. Analyses were done in R version 4.2.1, using the packages ggplot2, glmmTMB, multcomp, wesanderson, terra.

anthophila

display_name: anthophila

cities

display_name: cities

dominance

display_name: dominance

sads

display_name: sads

urban-beekeeping

display_name: urban-beekeeping

wildbees

display_name: wildbees

en: Data supporting ‘Examining honeybee (Apis mellifera) dominance patterns within urban bee communities worldwide’


en: field-data well-being artificial-lighting urban-landscape stress

en: 05_Anonymised dataset.xlsx

en: This file contains data for the manuscript "Randomised trial reveals a mismatch between preferences for and hormonal responses to anthropogenic light colour temperatures".

en: 02_Particiants consent form.pdf

en: This file contains the informed consent document that participants signed to participate in the study.

en: This document includes the questionnaire administered at the first measurement time point (before exposure).

en: This document includes the questionnaire administered at the second measurement time point (after exposure).

en: 01_Information sheet.pdf

en: This file includes the information sheet provided to participants before they consented to take part in the study.

en:
This repository contains data related to the field experiment of the WP3 (social impacts) of the ALANeX project. This project is a collaboration between the Swiss Federal Institute for Forest, Snow and Landscape Research (WSL) and Elektrizitätswerk des Kantons Zürich (EKZ). It has received funding from the WSL programme EXtremes. The overall objective of the ALANeX project was to assess the ecological and social impact of anthropogenic light at night.
We conducted a randomized field experiment to compare the well-being benefits of exposure to streetlights of different correlated colour temperatures (CCT) during a 20-min stay in the centre of Richterswil, Swizerland, to explore:
 - the impact of exposure to streetlamps of different CCT on affect, self-reported stress and cortisol levels
- the differences in impression of artificial light and of the feeling of safety provided by streetlamps based on its CCT.

artificial-lighting

display_name: artificial-lighting

field-data

display_name: field-data

stress

display_name: stress

urban-landscape

display_name: urban-landscape

well-being

display_name: well-being


en: photogrammetry avalanches davos dischma snow-hydrology snow-depth-distribution

en: Snowdepth maps

en:
The available datasets are snow depth maps with a spatial resolution of 0.5 m derived from images of the survey camera Vexcel Ultracam mounted on a piloted airplane. Image acquisition was carried out during the approximately peak of winter (time when the thickest snowpack is expected) in spring. The snow depth maps are calculated by the subtraction of a summer-DTM from the processed winter- DSM of the corresponding date. The summer-DTM used was derived from a point cloud of an airborne laser scanner from 2020.

Due to the occurrence of inaccuracies of the calculated snow depth values caused by the photogrammetric method, we applied different masks to significantly increase the reliability of the snow depth maps. We masked out settled areas, high-frequented streets and technical constructions, pixels with high vegetation (height > 0.5 m) , outliers and unrealistic snow depth values. In addition, we modified the snow depth values of snow-free pixels to 0.
The information on buildings and infrastructure comes from the exactly classified ALS point cloud and the TLM dataset from Swisstopo (https://www.swisstopo.admin.ch/de/geodata/landscape/tlm3d.html#links). High vegetation is also derived from the classification and the calculated object height from the point cloud.
Outliers and unrealistic snow depth values are defined as negative snow depth values and snow depths exceeding 10 m. The classification of each pixel of the corresponding orthophoto into snow-covered or snow-free is based on the application of a threshold of the NDSI or manually determined ratios of the RGB values.

An extensive accuracy assessment proves the high accuracy of the snow depth maps with a root mean square error of 0.25 m for the year 2017 and 0.15 m for the following snow depth maps.

The work is published in:

avalanches

display_name: avalanches

davos

display_name: davos

dischma

display_name: dischma

photogrammetry

display_name: photogrammetry

snow-depth-distribution

display_name: snow-depth-distribution

snow-hydrology

display_name: snow-hydrology

educ

en: Education, culture and sport

envidat

en: EnviDat: the Environmental Data Portal

en: EnviDat is the WSL data portal providing unified and managed access to environmental monitoring and research data. The portal has the capability to host and publish data sets. Through the sharing of unique and high-value data sets worldwide, EnviDat fosters research and contributes to advancing environmental science.

en: Snow depth mapping by airplane photogrammetry (2017 -  ongoing)


snow-drift-station-flowcapt

maintainer: Benjamin Walter

title_for_slug: snow-drift-station-flowcapt

en: raclets acoustic-mass-flux snow-blowing snowdrift flowcapt wind-speed

en: avergaged mass flux

en: Average mass flux of 10 min measurement interval. The Cycle duration is 1 min which includes 6 s observation and 54 s stand-by. The time is given in UTC+1.

en: minimum mass flux

en: The data file contains the minimum mass flux of 10 minute intervalls. The duration of one cycle was 1 min and contained 6s observations and 54 s stand-by. The time is given in UTC+1.

en: maximum mass flux

en: The data file contains the maximum mass flux of 10 minute intervalls. The duration of one cycle was 1 min and contained 6s observations and 54 s stand-by. The time is given in UTC+1.

en: minimum wind spees

en: The data file contains the minimum wind speed in 10 minute intervalls. The duration of one cycle was 1 min and contained 6s observations and 54 s stand-by. The time is given in UTC+1.

en: average wind speed

en: The data file contains the average wind speed in 10 minute intervalls. The duration of one cycle was 1 min and contained 6s observations and 54 s stand-by. The time is given in UTC+1.

en: maximum wind speed

en: The data file contains the maximum wind speed in 10 minute intervalls. The duration of one cycle was 1 min and contained 6 s observations and 54 s stand-by. The time is given in UTC+1.

en: The FlowCapt is an ultra-robust instrument measuring solid particle acoustic mass - flux intensities (g m‾ ² s‾ ¹) and wind speeds (m s‾ ¹). It was deployed at Gotschnagrat (LON: 46.859 LAT: 9.849). The vertical tube with a length of 1 m monitors snowdrift and snow-blowing; and is mounted at a height between 0.1 an 1.1 m above snow surface. The time within each data set is given in UTC+1.

acoustic-mass-flux

display_name: acoustic-mass-flux

flowcapt

display_name: flowcapt

raclets

display_name: raclets

snow-blowing

display_name: snow-blowing

snowdrift

display_name: snowdrift

wind-speed

display_name: wind-speed

en: Snow Drift Station - Flowcapt